Novalena lutzi (Gertsch 1933)

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 25-27

Novalena lutzi (Gertsch 1933)

Figure 12 ; Map 2

Agelena lutzi Gertsch 1933: 12 , figure 9 (♂)

Agelena idahoana Gertsch 1934: 25 , figure 5 (♀) NEW SYNONYMY Novalena idahoana Chamberlin & Ivie 1942: 229 , figure 57 (♀)

Novalena lutzi Chamberlin & Ivie 1942: 229 , figures 59–60 (♂ ♀) Novalena wawona Chamberlin & Ivie 1942: 230 , figure 58 (♀) NEW SYNONYMY

Types. Holotype male of N. lutzi : USA: Colorado: Mesa Verde Park , 37°N, 108°W, 2103–2347 m, 3–7.VII.1919, F.E. Lutz, deposited at AMNH (examined) . Holotype female of N. idahoana : USA: Idaho: Montpelier , 42°18′N, 111°18′W, VIII.1931, W.J. Gertsch, deposited at AMNH (examined) . Holotype female of N. wawona : USA: California, Yosemite Park, Wawona Camp , 37°N, 119°W, 17.IX.1941, W. Ivie, deposited at AMNH (examined).

Synonymy justification. Only the females of N. idahoana and N. wawona are described. They show the same pattern of the females identified as N. lutzi : shape of the atrium in posterior view, median plate uniformly wide and copulatory openings wider than long as illustrated by Gertsch (1934: fig. 5). The differences in Chamberlin and Ivie’s illustrations (1942: figs. 57, 58, 60) are probably due to different orientations in the drawings and the slightly damaged condition of the epigynum of N. idahoana .

Diagnosis. The males of N. lutzi distinguish from N. calavera in having the RTA with less pointed tips on dorsal projection ( Fig. 12 b) and ventral projection more prominent ( Fig. 12 a), and from N. rothi and N. alamo in having the tegular median process less conspicuous ( Fig. 12 a). The females differ from N. calavera in having the median plate uniformly wide and the copulatory openings wider than long ( Fig. 12 e), and from N. rothi in having hoods and the copulatory openings not visible in ventral view ( Fig. 12 c).

Description. Male (holotype) of N. lutzi : Carapace light brown. Chelicerae and condyles dark brown. Endites and labium brown with white tips. Sternum brown. Legs brown. Three rings on femur, one on patella, and two on tibia. Opisthosoma brown with several black spots and light brown foliage. Spinnerets orange, PLS with basal segment dark brown. Total length 5.66. Carapace length 2.93, width 2.33, cephalic region width 1.9, ocular region width 1. Chelicerae with two retromarginal teeth; basal segment length 1.4, fang length 0.65. Labium as long as wide (0.5/0.5). Endites convergent (distance at their base compared at their tips 0.5/0.2). Sternum longer than wide (1.8/1.3). Opisthosoma longer than wide (2.66/2). ALS separated by less their basal diameter (0.2/0.3), PLS with distal segment shorter than basal segment (0.35/0.5). Leg lengths: I—femur 3.7/ patella-tibia 4.5/ metatarsus 3.7/ tarsus?; II—3.5/ 4.6/ 4/ 2; III—3.4/ 4.2/ 4.1/ 1.9; IV—3.5/?/?/?. Spination: femur dorsal I—1-3 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-3 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—?/?/?; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 1-1-0; II—0/ 2-2-2/ 1-1-0/ 1-1-0; III—1-1 -0/ 2-2-2/ 1- 1-0/ 1-1-0; IV—?/?/?/?; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-1-0; II—0/ 2-2-2/ 0-1-0/ 0-1-0; III—3-2 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—?/?/?/?. Number of trichobothria on tarsus: I—?, II—6, III—5, IV—?. Number of dorsal spines on pedipalp: femur 2, tibia 3 Cymbium length 1.5, width 0.8. Embolus curved; conductor with dorsal projection larger than ventral one; tegular median process slightly sclerotized; dorsal projection of RTA with two sharp tips and a deep excavation, ventral projection prominent ( Fig. 12 a, b).

Female (holotype) of N. wawona : Carapace yellow. Chelicerae dark brown, condyles orange. Endites and labium brown with white tips. Sternum brown. Legs brown with three rings on femur and two on tibia. Opisthosoma dark brown with reddish foliage. Spinnerets light brown. Total length 11. Carapace length 5.4, width 3.7, cephalic region width 2, ocular region width 1.1. Chelicerae with two retromarginal teeth; basal segment length 1.8, fang length 0.9. Labium slightly longer than wide (0.8/0.7). Endites slightly convergent (distance at their base compared at their tips 0.7/0.4). Sternum longer than wide (2.8/2.2). Opisthosoma longer than wide (6.2/ 3.8). ALS separated by less their basal diameter (0.3/0.4), PLS with distal segment as long as basal segment (0.6/ 0.6). Leg lengths: I—femur 4.2/ patella-tibia 5.2/ metatarsus 3.8/ tarsus 2.2; II—4.2/ 4.8/ 3.8/ 2; III—3.9/ 4.8/ 4.1/ 1.9; IV—4.85/ 6/ 5.5/ 2.3. Spination: femur dorsal I—1-2 -3/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -2; patella I—dorsal 1- 1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 1-1-1-2/ 1-1-0/ 1-1-0; metatarsus I—0-0-1/ 2-2-2/ 0-1-1/ 0-0-1; II—0/ 2-2-2/ 0-1-1/ 0-1-1; III—2-2 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-2 -2/ 1- 1-1-2-2/ 0-1-1/ 0-0-1. Number of trichobothria on tarsus: I—7, II—7, III—7, IV—7. Dorsal spines on pedipalp femur: 3. Epigynal plate wider than long (2.5/1); atrium divided, visible in posterior view; anterior hoods present; copulatory openings wider than long ( Fig. 12 c, e). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent and separated from posterior margin ( Fig. 12 d).

Variation. Males (n=8): body length 5.66–7.88; carapace length 2.93–4; patella-tibia I length 4–4.75. Females (n=11): body length 8.75–13; carapace length 4–5.4; patella-tibia I length 4.13–5.2. One female with two to three retromarginal teeth on chelicerae.

Additional material examined. USA: Arizona: Pinecrest, Graham Mountain , 13.IX.1950, W.J. Gertsch, 1 ♀ ( AMNH) ; same data, 1 ♀ ( AMNH) ; Jacob Lake , 36°N, 112°W, 14.VI.1934, Ivie and Rasmussen, 2 ♂ ( AMNH) ; Santa Catalina Mountains , 30.X.1937, Bryant, 1 ♀ ( AMNH) . California: El Dorado Co., 6.4 km W Kyburz, 15.IX.1959, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH) ; Mariposa Co., 1.6 km S Fish Camp, 23.IX.1961, W. Ivie and W.J. Gertsch, 1 ♀ ( AMNH) ; Tulare Co., Cedar Grove, Kings River Canyon , 16.VII.1952, M. Cazier, W.J. Gertsch, and R. Schrammel, 2 ♀ ( AMNH) ; Tulare Co. 1.6 km S Belnap Springs near Nelsons Camp , 11.VII.1958, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH) ; Tulare Co., Moorehouse Creek , near Nelson Camp, 11.VII.1958, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH) ; Tulare Co., Sequoia National Park , 11.3 km NE Ash Mountain, 14.IX.1959, W.J. Gertsch and V. Roth, 1 ♀ ( AMNH) ; Sequoia National Park near Ash Mountain , 9.VII.1958, W.J. Gertsch and V. Roth, 4 ♀ ( AMNH) ; Road’s End Kern River , 3.VII.1956, V. Roth and W.J. Gertsch, 1 ♀ ( AMNH) ; Yosemite National Park, Glacier Point Road , 3–4.VII.1946, S. Mulaik and D. Mulaik, 1 ♀ ( AMNH) . Colorado: Gunnison Co., East Portal Black Canyon , 1996 m, 26.VII.1962, W.J. Gertsch, 4 ♀ ( AMNH) ; Routt Co., Hot Springs , 26.IX.1983, V. Roth and B. Roth, 1 ♀ ( AMNH) ; Pike National Forest , 38°N, 105°W, 4.IX.1959, D.T. Jennings, 1 ♂ ( AMNH) ; 24.1 km W Denver , 24.VIII.1935, W. Ivie, 1 ♀ ( AMNH) ; San Juan National Forest, Lower Piedra Campground , 13.VI.1946, no collector data, 1 ♀ ( AMNH) . Nevada: Charleston Mountain , 2438 m, 6.VIII.1944, D.J. Zinn, 1 ♂ ( AMNH) . New Mexico: 1.6 km S Capillo Peak , NW Manzano, no date data, C.C. Hoff, 1 ♀ ( AMNH) ; Bernalillo Co., Sandia Mountains , no date data, C.C. Hoff, 1 ♀ ( AMNH) . Utah: American Fork Canyon , 40°N, 111°W, 20.VIII.1927, R.V. Chamberlin, 1 ♀ ( AMNH) ; Cedar Canyon , 37°N, 112°W, 10.VI.1927, R.V. Chamberlin, 1 ♀ ( AMNH) ; City Creek Canyon, Rotary Grove , 40°48′N, 111°48′W, 22.IX.1946, W. Ivie, 3 ♀ ( AMNH) ; same data except VIII–IX.1947, 3 ♀ ( AMNH) ; same data except 40°N, 111°W, 11.IX.1947, 1 ♀ ( AMNH) ; Henry Mountains, Horse Valley , 7.IX.1929, Chamberlin and Gertsch, 2 ♀ ( AMNH) ; Mill Creek Canyon, 40°N, 111°W, 18.VIII.1941, J.C. Chamberlin, 1 ♀ ( AMNH) ; Price , 39°N, 110°W, IX.1940, H. Higgins, 1 ♀ ( AMNH) .

Distribution. USA: Arizona, California, Colorado, Idaho, Nevada, New Mexico, Utah (Map 2).

Figures

FIGURE 12. Novalena lutzi (Gertsch 1933): a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; CO, copulatory openings; E, embolus; FD, fertilization ducts; H, hoods; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: 0.5 mm.