Thecomyia lateralis ( Walker, 1858 )

Marinoni, Luciane, Steyskal, George C. & Knutson, Lloyd, 2003, Revision and cladistic analysis of the Neotropical Genus Thecomyia Perty (Diptera: Sciomyzidae), Zootaxa 191, pp. 1-36 : 16-17

2. Thecomyia lateralis ( Walker, 1858) ( Figs. 7, 19 , 31, 41, 44)

Tetanocera lateralis Walker, 1858: 221 [type­locality: "Valley of Amazon"]. Thecomyia lateralis .­ Steyskal, 1966: 447 [type examination, generic combination].­ Knutson et al., 1976: 13 [Neotropical catalog].

Thecomyia trilineata Hendel, 1932: 98 [type­locality: Pozo del Tigre, Bolivia]. New Synonym.

Description. Adult. Upper orbital seta absent. Fronto­orbital spot rounded, width approximately 1/3 of frontal width. Rostrum 0.66­0.72 in males and 0.71­0.78 in females as high as eye. White microtomentose infra­ocular marking forming a triangle with margin between gena and postgena emarginate. Mid femur without anteromedial seta. Mid and hind femora usually wholly yellowish. Mesonotum with ill­defined medial light stripe, sometimes with 3 lighter lines; scutellum yellow. Ejaculatory apodeme very broad at apex ( Fig. 7). Phallapodeme slightly concave at articulation with basiphallus, without subapical process; slightly curved. Anterior basiphallic process with apex bifurcated. Epiphallus long, well developed, extending to base of distiphallus. Distiphallus with acrophallus coiled. Hypandrium in lateral view, with dorsal margin almost straight; anteroventral margin narrow, rounded ( Fig. 19 ). Anteroventral hypandrial plates in frontal view very narrow, tapered laterally. Membranous projections of margins of anteroventral hypandrial plates well developed, not extending to lateral sides of hypandrium, covered with minute hairs ( Fig. 19 ). Anterior surstylus absent, posterior appearing as a bar ( Fig. 7). Female sternites 6, 7 and 8 almost fused, longer than wide; anterior margin of sternite 6 slightly sinuate; mid longitudinal portion almost membranous, with very weak sclerotization, sometimes folded. Spermathecal duct with sclerotized region joining spermatheca long; apical digitiform projection thin, short (Fig. 31).

Type specimens. Tetanocera lateralis: Condition generally poor, head and thorax greasy, with remnants of mold; left wing destroyed in apical and posterior 1/2; abdomen missing. There are five labels as follows: 1. White circular label with green border, “ Type ” in black print; 2. “ Tetanocera lateralis Walk ” in black ink on white paper; 3. “S. America Brazil ” in black ink on white paper, same hand as in number 2; 4. “ lateralis ” in black ink on white paper folded three times (different hand than numbers 2 and 3); 5. “ Brazil ” in black ink on white paper (same hand as label 4). Comments on the type specimens.­L. Knutson (19 Nov 1998) examined the male type of Tetanocera lateralis in the Natural History Museum, London, and noted that the abdomen was missing. At that time John E. Chainey searched the collection, including the slide collection, for the missing parts but without success. According to correspondence of 29 Apr 1968, Brian H. Cogan prepared and sent a drawing of the aedeagus of T. lateralis to G. C. Steyskal.

Thecomyia trilineata: In good condition except for the missing antennae, left fore tarsus and right mid leg; left mid leg glued to the label; genitalia in microvial in glycerin. There are three labels as follows: 1. White rectangular label with black border, "Pozo del Tigre Boliv." "Lindner, D. Chaco ­ Exped." in black handwriting; 2. White rectangular label with black border, " Thecomyia trilineata Hendel " in black handwriting. 3. White rectangular label with black border, " Type Hendel 1952" in red handwriting. The male type of T. trilineata was lent by the Staatliches Museum für Naturkunde in Stuttgart and examined by L. Marinoni.

Other specimens examined. GUYANA. Mus. Exp. BE Dahligren, no date, no col., 1 m. ( FMNH); 1908, K.S. Wise col., 1 m. and 1 f. ( BMNH); Kanuku Mts, Moco Moco River, 3o 18. 2'N, 59o 38. 9'W, 29 Apr 1995, W. N. Mathis, 1 m ( USNM). SURINAME. Anapaike (Rio Lawa), Marowijne, Nov 1963, B. Malkin col., 1 f. ( MZSP). BRAZIL. AMAZONAS: Benjamin Constant, 18­28 Nov 1962, K. Lenko col., 1 f. ( MZSP); BAHIA: Itabuna, Fazenda Bela Vista, Jan 1964, no col., 1 m. ( MZSP); MINAS GERAIS: Andes, Feb 1956, M. Carrera col., 1 f. ( MZSP); MATO GROSSO: Maracaju, Feb 1937, no col., 2 m. ( USNM); GOIÁS: Corumbá, Fazenda Monjolinho, Nov 1945, Barreto col., 1 m. ( MZSP); Jataí, Jan 1955, M. Carrera, A. Machado, F.S. Pereira & E. D. M. Loureiro cols., 1 f. and 1 no abdomen ( MZSP); RIO DE JANEIRO: no locality, Jul 1908, W. M. Wheeler Collection, no col., 1 m. ( AMNH), SÃO PAULO: Tamoio, Rio Jacaré, Dec 1944, Barreto col., 1 f. ( MZSP); Magda, Dec 1956, J. Lane col., 1 f. ( MZSP); Est. Biol. Boracéia, Salesópolis, 850 m., 2 May 1963, Rabello col., 1 f. ( MZSP); Iporanga, 1 Nov 1961, Lenko & Reichardt, 1 f. ( MZSP); Avanhadava, Barra Mansa, Feb 1946, Barreto col., 1 m. and 1 f. ( MZSP); Estrada Rio­São Paulo, 19 Feb 1946, L. Miranda col., 1 m. ( MZSP); Araçatuba, Córrego Azul, Mar 1947, M. P. Barreto col., 5 m. and 6 f. ( MZSP); same locality, Sep. 1946, Barreto col., 2 m. ( MZSP); Rio Jacarecatinga, Oct 1961, Lane & Rabello cols., 1 m. and 1 f. ( MZSP); Ilha dos Búzios, 16 Oct­ 4 Nov 1963, Exp. Dep. Zool., 1 m. ( MZSP); PARANÁ: Morretes, Sapitanduva, 9 Nov 1986, L. Marinoni, C. Carvalho & L. Knutson cols., 1 m. ( USNM); same locality, same date, same cols., 2 f. ( DZUP); SANTA CATARINA: Florianópolis, Jul 1960, Casemiro col., 2 m. and 1 f. ( MZSP). PERU. Iquitos, Mar­Apr 1931, R. C. Shannon, 6 m. and 4 f. ( USNM); Loreto, Ucayali River, Yarina Cocha , 22 Nov 1953, Peter Hocking col. 1 f. ( FMNH). BOLIVIA. Andrés Ibañez, Potrerilo del Guenda, 12 Sep 1993, P. Bettella col., 1 m. ( BMNH).

Figures

FIGURES 18 ­ 23. Hypandrium, frontal view and right lateral view: 18, Thecomyia longicornis Perty; 19, T. lateralis (Walker); 20, T. limbata (Wiedemann); 21, T. abercrombiei Marinoni and Steyskal; 22, T. autazensis Marinoni and Steyskal; 23, T. bonattoi Marinoni and Steyskal. A, anterior; as, archlike struts; avp, anteroventral plates; d, dorsal; mp, membranous projection; v, ventral.