Trocniassus

Dai, Wu, Dietrich, Christopher H. & Zhang, Yalin, 2015, A review of the leafhopper tribe Hyalojassini (Hemiptera: Cicadellidae: Iassinae) with description of new taxa, Zootaxa 3911 (1), pp. 1-142 : 36-37

Trocniassus gen nov.

Type species: T. shaanxiensis sp. nov.

Stramineous to brown, without darker spots or other distinct markings ( Fig. 5 E–H). Body robust, cylindrical, not or only weakly depressed. Head in dorsal view narrower than pronotum ( Fig. 5 E, G); crown weakly developed, very short in dorsal view; vertex nearly vertical, vertex and frontoclypeus transversely rugose; face distinctly broader than tall; gena obtusely emarginated below eye; rostrum slender, extended slightly beyond front trochanter; anteclypeus weakly convex, approximately parallel-sided, apex truncate, even with or not extended to lower margin of gena; lorum narrow, well separated from lateral and ventral margins of gena, with basal fourth of mesal margin bordering frontoclypeus; frontoclypeus convex; antennal ledge well developed, broad, slightly oblique; lateral frontal suture obsolete or extending only part way from antennal ledge to ocellus; ocellus large, directly above antennal pit, approximately half way between eye and midline and near mid height of vertex. Pronotum strongly convex, declivous anteriorly, transversely rugose; lateral margins long, strongly carinate, evenly divergent posterad ( Fig. 5 E, G). Exposed part of mesonotum and scutellum shorter than pronotum, mesonotum granulose, scutellum transversely rugose; scutellar suture arched anterad, scutellum weakly convex with acuminate apex. Forewing venation moderately well delimited; membrane translucent, without opaque sclerotization, punctate, without conspicuous setae; appendix broad, extended to wing apex, slightly narrower than inner apical cell; vein R with 8–14 branches; three closed anteapical cells present; apical cell 1 elongate, narrow, extended to wing apex; apical cell 2 much shorter, strongly narrowed distally; apical cells 3–4 short, parallel-sided, variable in length; posterior branch of CuA connected to margin at apex of clavus; clavus truncate apically; texture of inner apical cell same as that of appendix but different from that of adjacent apical cell ( Fig. 5 E–H). Front femur with pair of dorsoapical macrosetae well developed; AM1 poorly differentiated; PD, AD and AM without well differentiated preapical setae; intercalary group consisting of irregular band of fine short setae; AV and PV each with 3–4 well differentiated macrosetae in basal half; tibia with dorsal surface flat and bicarinate, AD and PD each with 7 macrosetae. Mesothoracic femur PD with 1 and AD with several macrosetae; AV with several macrosetae. Hind femur macrosetae 2+2+1 with antepenultimate seta as large as others; tibial rows PD, AD, and AV with 19, 13, and 18 macrosetae, respectively; tarsomere I without dorsoapical setae, PV with row of 4 stout cucullate setae continuous with 2 similar setae on tarsomere II, AV with scattered setae, pecten with 6 platellae.

Male abdominal apodemes poorly developed. Sternite VIII slightly longer than sternite VII, about half as long as broad, lateral margins parallel, apex convexly produced. Genital capsule with posteroventral part mostly concealed by sternite VIII in repose, with only anal tube and part of posterior pygofer margin visible. Pygofer with tergite long, well sclerotized, narrowly emarginate distally in dorsal view; lobes not elongated, rounded distally with band of macrosetae extended along most of posterior margin, fine setae present or absent; submarginal ridge present and extended from base of lobe to or beyond midheight; posterior margin concave, without processes; anterodorsal margin in lateral view emarginate along intersegmental membrane; intersegmental membrane between anal tube and aedeagal atrium without accessory lobes, sclerites, or setae; basolateral setal group absent (Fig. 21A, 22A–C). Subgenital plates articulated to valve, separated at base, convergent distally, moderately wide at base and narrowed distally, apex curved laterad with darkly pigmented ventral fold or projection, few scattered fine setae present (Fig. 21C, 22E). Valve moderately long, subquadrate, broadly fused to pygofer at base, posterior margin concave (Fig. 21A, 22A). Connective well sclerotized, Y-shaped, nearly vertical articulated to anteroventral corner of aedeagal atrium, anterior arms articulated styles (Fig. 21E, 21F, 22D, 22F). Style well developed, articulated to connective well below aedeagal atrium, in lateral view with apodeme nearly straight; apophysis longer than apodeme, curved dorsad or arcuate, apex acuminate, preapical projections absent (Fig. 21D, 22D). Aedeagus simple, short, atrium large with basolateral dorsal arms strongly divergent, ventrolateral projections absent; shaft tubular basally, with dorsal ridge distally and pair of ventrolateral longitudinal ridges preapically, curved dorsad in lateral view, apex entire, without processes; gonopore preapical on posterior surface (Fig. 21E, 21F, 22D, 22F).

Female sternite VII truncate; first valvulae moderately slender, evenly curved, dorsal sculpturing imbricate, apex abruptly tapered and slightly attenuated ( Fig. 24 C, D); second valvulae moderately slender, dorsal margin with four widely spaced teeth in distal half, apex evenly tapered to bluntly rounded tip, finely serrate dorsally ( Fig. 24 A, B).

Notes. Species of this genus resemble Trocnadella externally but differ in having a distinct row of cucullate setae on hind tarsomeres I and II and in having the connective of the male sclerotized and articulated to the aedeagus. The genus name, a masculine noun, was formed by combining Trocnadella with Iassus , the names of two previously described iassine genera.

Figures

FIGURE 5. Dorsal and lateral habitus. A, B, Lamelliassus chiangmaiensis sp. nov.; C, D, Siamiassus constanti sp. nov.; E, F, Trocniassus shaanxiensis sp. nov.; G, H, Trocniassus henanensis sp. nov.

FIGURE 24. Trocniassus shaanxiensis sp. nov. female genitalia. A. Apical half of second valvulae. B. Second valvulae of female ovipositor, lateral view; C. Apical half of first valvulae; D. First valvulae of female ovipositor, lateral view.