Panjange thomi Huber, 2019

Huber, Bernhard A. & Carvalho, Leonardo S., 2019, Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae), Zootaxa 4546 (1), pp. 1-96 : 80-81

Panjange thomi Huber sp. n.

Figs 300–303 , 308–311 , 316–318 , 333–335

Panjange Ind 103: Eberle et al. 2018 (molecular data); Huber et al. 2018: fig. 10.

Note. Females of this species were compared directly with the female holotype of Panjange dubia ( Kulczyński, 1911) : the epigynum is indistinguishable externally. However, several species in the Panjange cavicola group have very similar female external genitalia (e.g., P. alba Deeleman-Reinhold & Deeleman, 1983 ; P. madang Huber, 2011 ), and the dark mark in the ocular area is smaller in all females below than in the P. dubia holotype. Thus, the specimens below are not considered to be P. dubia even though they originate from closer to the type locality of P. dubia than the male tentatively described as P. dubia in Huber (2011a) (175 km vs. 400 km; Fig. 350 ).

Type material. INDONESIA: ♂ holotype, ZFMK (Ar 20660), West Papua, Manokwari, Gunung Meja (0.860°S, 134.084°E), 190 m a.s.l., 8.xi.2009 (S. Sutono) .

Other material examined. INDONESIA: 1♂ 9♀ 2 juvs, ZFMK (Ar 20661–62), and 2♀ 4 juvs in pure ethanol, ZFMK (Ind199), same data as holotype .

Etymology. The species is named for Thomas Wapai N.S. Wainggai (also known as Dr. Thom) (1937–1996), West Papuan scholar and political leader.

Diagnosis. Males can be distinguished from other representatives of the cavicola group with strongly elongated palps ( P. alba , P. dubia , P. madang , P. mirabilis ) by shape of procursus ( Figs 308–309 ; only one ventral ridge, distinctive flat distal elements) and by extremely elongated male palpal tibia (Figs; 308–309> 7 x longer than wide; other species: <5 x longer than wide). Females of these five species may be indistinguishable externally, but internal genitalia of P. thomi with distinctive structures diverging towards anterior ( Figs 311 , 318 ).

Description. Male (holotype). MEASUREMENTS. Total length 3.7, carapace width 0.87. Distance PME-PME 400 µm; diameter PME 90 µm; distance PME-ALE 40 µm; AME absent. Leg 1: 38.2 (8.8 + 0.4 + 9.0 + 18.1 + 1.9), tibia 2: 5.8, tibia 3: 3.2, tibia 4: 4.7; tibia 1 L/d: 113.

COLOR (in ethanol). Carapace pale whitish with dark median mark posteriorly and dark brown mark in ocular area including eye stalks; clypeus not darkened; sternum whitish; legs ochre-yellow, with brown rings in patella area and at tibia-metatarsus joints. Abdomen pale gray, with dark marks dorsally and laterally.

BODY. Habitus as in Fig. 300 . Eye triads on stalks ( Fig. 310 ), each stalk with single pointed process curved towards anterior. Thoracic furrow absent. Clypeus unmodified. Sternum wider than long (0.54/0.44), unmodified. Abdomen cylindrical, angular dorso-posteriorly.

CHELICERAE. As in Fig. 310 , with pair of simple weakly sclerotized processes proximally.

PALPS. As in Figs 308–309 , symmetric; coxa with small but distinct retrolateral-ventral apophysis, trochanter with retrolateral-dorsal process, femur with rounded prolateral process proximally; femur-patella joints strongly shifted towards prolateral side; tibia strongly elongated; tarsus with long whitish dorsal process with tarsal organ at tip; procursus with only one ventral ridge, without ventral pointed process, with distinctive distal flat elements; genital bulb oval, with two very long processes lying parallel to each other: semitransparent embolus and slightly sclerotized bulbal apophysis (‘appendix’; ba in Fig. 308 ).

LEGS. Without spines and curved hairs, few vertical hairs; retrolateral trichobothrium of tibia 1 at 3%; tibia 1 without prolateral trichobothrium (present on other tibiae); tarsus 1 with ~30 pseudosegments, only distally a few fairly distinct.

Male (variation). Tibia 1 in other male: 9.1.

Female. In general similar to male ( Figs 302–303 ) but eye triads on low humps, closer together (distance PME-PME 210 µm) and dark mark in ocular area smaller (only dark band between eye triads). Tibia 1 in 8 females: 7.6–9.0 (mean 8.2). Tarsus 4 comb-hairs as in Fig. 334 . Epigynum weakly sclerotized ( Fig. 316 ), with long folded scape ending in rounded ‘knob’ ( Figs 317 , 333 ). Internal genitalia with pair of roundish pore plates and distinctive structures diverging towards anterior ( Figs 311 , 318 ). ALS with one widened, one pointed, and six cylindrically shaped spigots ( Fig. 335 ).

Distribution. Known from type locality only ( Fig. 350 ).

Natural history. The spiders were collected from the undersides of green leaves.

Figures

FIGURE 350. Known distribution of Panjange in eastern Indonesia (not shown: record of P. alba from Ambon; see Huber 2011, fig. 435).

FIGURES 300–307. Panjange spp. live specimens. 300–303. P. thomi sp. n., adult and penultimate instar males, and female with egg sac, from Gunung Meja. 304–307. P. togutil sp. n., adult and penultimate instar males, females with and without eggsac, from near Sofifi.

FIGURES 308–309. Panjange thomi sp. n. (ZFMK, Ar 20661), left male palp, prolateral and retrolateral views; arrow points at single ventral ridge on procursus. ba, bulbal apophysis; em, embolus; gb, genital bulb; pr, procursus; tp, tarsal process. Scale line: 1.0.

FIGURES 310–311. Panjange thomi sp. n. (ZFMK, Ar 20661–62), male prosoma and chelicerae, oblique frontal view, and cleared female genitalia, dorsal view. Scale lines: 0.3.

FIGURES 316–321. Panjange spp., epigyna in ventral views, and cleared female genitalia in ventral and dorsal views. 316– 318. P. thomi sp. n. (ZFMK, Ar 20662). 319–321. P. togutil sp. n. (ZFMK, Ar 20665). Scale lines: 0.3.

FIGURES 329–335. Saciperere catuaba sp. n. (329–332, ZFMK, Ar 20635) and Panjange thomi sp. n. (333–335, ZFMK, Ar 20662). 329. Male gonopore. 330. Female palpal tibia. 331–332. Male and female spinnerets. 333. Epigynum, ventral view. 334. Female tarsus 4 comb hairs. 335. Female ALS. ALS, anterior lateral spinneret; PMS, posterior median spinneret. Scale lines: 10 µm (329, 331-332, 334-335), 20 µm (330), 100 µm (333).