Anthobosca Guerin de Meneville

Kimsey, Lynn S., 2011, Tiphiidae wasps of Madagascar (Hymenoptera, Tiphiidae), Journal of Hymenoptera Research 22, pp. 45-68 : 47-49

Genus Anthobosca Guerin de Meneville Fig. 2

Anthobosca Guérin de Méneville 1838:237.

Type species:

Anthobosca australasiae Guérin de Méneville 1838:237.

Discussion.

This is the least derived genus in the family Tiphiidae ( Kimsey 1991). Both sexes are fully winged, with complete venation. Males are slenderer than females, with linear, cylindrical antennae. Females are less commonly collected than males, and are heavy-bodied with coiled antennae. The face is flat, without supraantennal lobes, similar to the face in Tiphiinae . There are fourteen Anthobosca species described from mainland Africa and ten from Madagascar ( Kimsey 2009). Hosts are unknown for the subfamily.

Key to species of Anthobosca (modified from Kimsey 2009)

1	Metasoma 6-segmented; ten flagellomeres; females	2
-	Metasoma 7-segmented; eleven flagellomeres; males	4
2	Head, mesosoma and metasoma concolorous brown; wing membrane amber-tinted; body over 25 mm long	Anthobosca insularis F. Smith
-	Head, mesosoma and metasoma blackish or blackish and red, often with whitish spots; wing membrane untinted or tinted with brown to black; body length less than 15 mm long	3
3	Metasoma black to dark brown, terga II-III with whitish lateral spots	Anthobosca madecassa Krombein
-	Metasoma red, terga II-III without lateral spot	Anthobosca dimidiata Bartalucci
4	Metasomal sterna II and III with large spoon-like ventral projections; metasoma with extensive yellow markings	Anthobosca hallucigenia Kimsey
-	Metasomal sterna II and III unmodified, without projections; metasoma without yellow markings	5
5	Body length 17-20 mm; wings dark amber-tinted; flagellomeres without tyloids; tegula brown	Anthobosca insularis F. Smith
-	Body length 11 mm or less; wings untinted; three or more flagellomeres with tyloids; tegula white	6
6	Flagellomere XI less than 2.2 × as long as broad	7
-	Flagellomere XI more than 2.4 × as long as broad	10
7	Flagellomere VIII without tyloids; clypeus evenly convex with narrow ventrally facing flattened apical surface	Anthobosca micromeria Bartalucci
-	Flagellomere VIII with one tyloid; clypeus without ventrally facing flattened apical surface	8
8	Flagellomere VII with one small tyloid	Anthobosca nigrimacula Kimsey
-	Flagellomere VII without tyloids	9
9	Volsella broadest submedially, dorsal lobe oriented diagonally; paramere apex brown	Anthobosca castanea Kimsey
-	Volsella broadest dorsally, dorsal lobe oriented horizontally; paramere apex whitish	Anthobosca toliaraensis Kimsey
10	Paramere apex white; flagellum strongly bicolored brown to black dorsally, yellow ventrally; digitus extending diagonally to horizontally toward paramere in profile	11
-	Paramere apex brown; flagellum brown to black, may be slightly paler ventrally; digitus extending diagonally toward paramere in lateral view	12
11	Clypeus evenly convex medially, entirely white or less commonly bicolored, black and white; volsella broadest dorsally, digitus horizontally oriented in lateral view	Anthobosca mahajangaensis Kimsey
-	Clypeus flattened medially, entirely brown or bicolored brown and black; volsella narrowed dorsally, digitus oriented obliquely in lateral view	Anthobosca namorokaensis Kimsey
12	Flagellomere XI 2.6-3.0 × as long as broad; digitus diagonally to vertically oriented in lateral view	Anthobosca harinhalai Kimsey
-	Flagellomere XI 2.3-2.4 × as long as broad; digitus horizontally oriented in lateral view	Anthobosca fisheri Kimsey

Checklist of Anthobosca species

1. Anthobosca castanea Kimsey 2009:2. Holotype male: Madagascar: Mahajanga Prov., Parc National d’Ankarafantsika, Ampijoroa Station Forestiére, 40 km 306° 2nw Andranofasika (SAN FRANCISCO). Distribution: Mahajanga and Toliara Provinces.

2. Anthobosca dimidiata Bartalucci 2005:1082. Holotype female; Madagascar: Tuléar Berenti, 12 km nw Amboasary (LONDON). Distribution: Toliara Prov.

3. Anthobosca fisheri Kimsey 2009:5.Holotype male: Madagascar: Toliara Prov. Forêt de Mahavelo Isantoria River (SAN FRANCISCO). Distribution: Toliara, Toamasina, Antsiranana and Mahajanga Provinces.

4. Anthobosca hallucigenia Kimsey 2009:9. Holotype male: Madagascar: Toamasina Prov., Andasibe National Park (SAN FRANCISCO). Distribution: Antsiranana, Finanarantsoa and Toamasina Provinces.

5. Anthobosca harinhalai Kimsey 2009:11. Holotype male: Madagascar: Toliara Prov. Forêt de Mahavelo Isantoria River (SAN FRANCISCO). Distribution: Toliara and Tuléar Provinces.

6. Anthobosca insularis (Smith) 1879: 178. ( Myzine ). Holotype female; Madagascar (LONDON). Distribution: Ranomafana National Park and adjacent Fianarantsoa Prov.

7. Anthobosca madecassa Krombein 1949:52. Holotype female; Madagascar: Bekily (ITHACA). Distribution: Antsiranana and Toliara Provinces.

8. Anthobosca mahajangaensis Kimsey 2009:12. Holotype male: Madagascar: Mahajanga Prov., Parc National de Baie de Baly (SAN FRANCISCO). Distribution: Mahajanga Prov.

9. Anthobosca micromeria Bartalucci 2005:1083. Holotype male; Madagascar: Tuléar Morondava (LONDON). Distribution: Mahajanga Prov.

10. Anthobosca namorokaensis Kimsey 2009:15. Holotype male: Madagascar: Mahajanga Prov., Namoroka National Park, 16.9 km, 317° nw Vilanandro (SAN FRANCISCO). Distribution: Mahajanga Prov.

11. Anthobosca nigrimacula Kimsey 2009:16. Holotype male: Madagascar: Toliara Prov., Réserve Spéciale de Cap Sainte Marie (SAN FRANCISCO). Distribution: Toliara Prov.

12. Anthobosca toliaraensis Kimsey 2009:16. Holotype male: Madagascar: Toliara Prov., 12 km se Tuléar, (DAVIS). Distribution: Mahajanga and Toliara Provinces.

Figures

Figures 1 - 6. Lateral view of males. 1 Methocha 2 Anthobosca 3 Tiphia (uncus obscured by genital capsule shown in inset) 4 Mesa 5 Meria 6 Myzinella.