Trombiculinae Ewing, 1929

Stekolnikov, Alexandr A., 2021, A checklist of chigger mites (Acariformes: Trombiculidae) of Southeast Asia, Zootaxa 4913 (1), pp. 1-163 : 40-49

publication ID	https://doi.org/10.11646/zootaxa.4913.1.1
publication LSID	lsid:zoobank.org:pub:E3540F07-4CF6-4EF2-A533-64A2D49D68FF
DOI	https://doi.org/10.5281/zenodo.4461744
persistent identifier	https://treatment.plazi.org/id/E016879A-6627-2964-33F3-A90BFDB91C57
treatment provided by	Plazi (2021-01-19 08:10:39, last updated 2024-11-25 18:48:34)
scientific name	Trombiculinae Ewing, 1929
status

Treatment

Subfamily Trombiculinae Ewing, 1929

Tribe Schoengastiini Vercammen-Grandjean, 1960

Genus Ascoschoengastia Ewing, 1946

Ascoschoengastia audyi ( Womersley, 1952)

Ascoschoengastia calcar Nadchatram and Domrow, 1964

Ascoschoengastia canus Domrow, 1962

Ascoschoengastia ctenacarus Domrow, 1962

Ascoschoengastia daria (Traub and Audy, 1954)

Ascoschoengastia dentata Nadchatram and Domrow, 1964

Ascoschoengastia globosa Nadchatram and Domrow, 1964

Ascoschoengastia gressitti Nadchatram and Domrow, 1964

Ascoschoengastia indica (Hirst, 1915)

= Ascoschoengastia monteli André, 1954

= Trombicula muris Walch, 1922

Ascoschoengastia indicella (Traub and Audy, 1954)

Ascoschoengastia kittii Domrow and Nadchatram, 1964

Ascoshoengastia krishnani Nadchatram and Domrow, 1964

Ascoschoengastia leechi Domrow, 1962

Ascoschoengastia malayensis ( Gater, 1932)

Ascoschoengastia lorius ( Gunther, 1939)

Ascoschoengastia montana Yu, Yang and Chen, 1980

Ascoschoengastia ocellifera (Traub and Audy, 1954)

Ascoschoengastia octavia Domrow, 1962

Ascoschoengastia roluis (Traub and Audy, 1954)

Ascoschoengastia rousetti Goff, 1981

Ascoschoengastia sellnicki Audy and Womersley, 1957

Ascoschoengastia soekaboemiensis ( Takekawa, 1945)

Ascoschoengastia tafia Nadchatram and Domrow, 1964

Genus Bishoplinia Vercammen-Grandjean and Nadchatram, 1965

Bishoplinia wilsoni Vercammen-Grandjean and Nadchatram, 1965

Genus Cheladonta Lipovsky, Crossley and Loomis, 1955

Cheladonta gouldi Lakshana, 1969

Cheladonta neda Schluger et al., 1960

Genus Doloisia Oudemans, 1910

Subgenus Doloisia Oudemans, 1910

Doloisia (Doloisia) alata Schluger et al., 1961

Doloisia (Doloisia) fulminans Schluger et al., 1961

Doloisia (Doloisia) hooperi Domrow and Nadchatram, 1962

Doloisia (Doloisia) nasicola Domrow and Nadchatram, 1962

Subgenus Traubacarus Audy and Nadchatram, 1957

Doloisia (Traubacarus) alticola (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) brachypus (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) browningi (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) domrowi (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) gigantea ( Schluger et al., 1961)

= Doloisia outoensis Chen and Hsu, 1965

Doloisia (Traubacarus) hamata Domrow and Nadchatram, 1962

Doloisia (Traubacarus) harrisoni (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) intermedia (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) jadini (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) manipurensis (Radford, 1946)

Doloisia (Traubacarus) owenevansi (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) varmai (Audy and Nadchatram, 1957)

Doloisia (Traubacarus) vercammeni (Audy and Nadchatram, 1957)

Genus Farrellioides Vercammen-Grandjean, 1960

Farrellioides consuetum ( Womersley, 1952) , comb. nov.

Farrellioides nakatae (Nadchatram and Traub, 1964) , comb. nov.

Farrellioides striatum (Nadchatram and Traub, 1964) , comb. nov.

Genus Guntheria Womersley, 1939

Subgenus Phyllacarus Vercammen-Grandjean, 1967

Guntheria (Phyllacarus) bushlandi ( Philip, 1947) , comb. nov.

Guntheria (Phyllacarus) crinita ( Womersley, 1952)

Guntheria (Phyllacarus) mohri ( Womersley, 1952)

Guntheria (Phyllacarus) philippensis (Philip and Woodward, 1946)

Genus Helenicula Audy, 1954

Helenicula comata ( Womersley, 1952)

Helenicula consonensis Hadi and Carney 1977

Helenicula globularis ( Walch, 1927)

Helenicula kohlsi (Philip and Woodward, 1946)

= Helenicula hongkongensis Womersley, 1957

Helenicula lanius (Radford, 1946)

= Neoschoengastia covelli Radford, 1953

Helenicula longipoda Nadchatram and Traub, 1971

Helenicula mutabilis ( Gater, 1932)

Helenicula naresuani Stekolnikov, 2016

Helenicula pilosa ( Abonnenc and Taufflieb, 1957)

Helenicula scanloni Domrow and Nadchatram, 1964

Helenicula selvana Schluger et al., 1960

Helenicula signata ( Womersley, 1952)

Helenicula simena ( Hsu and Chen, 1957)

Helenicula ungkomari Nadchatram and Traub, 1971

Genus Herpetacarus Vercammen-Grandjean, 1960

Subgenus Abonnencia Vercammen-Grandjean, 1960

Herpetacarus (Abonnencia) cadigani Nadchatram and Kethley, 1974

Herpetacarus (Abonnencia) leprochaeta Nadchatram and Kethley, 1974

Subgenus Cricacarus Vercammen-Grandjean, 1966

Herpetacarus (Cricacarus) thompsoni (Domrow, 1962)

Genus Kayella Vercammen-Grandjean, 1960

Kayella masta ( Traub and Sundermeyer, 1950) , comb. nov.

Kayella novita ( Audy and Womersley, 1957)

Genus Mackiena Traub and Evans, 1950

Mackiena empodiformis Traub and Evans, 1950

Genus Neoschoengastia Ewing, 1929

Neoschoengastia americana ( Hirst, 1921)

= Neoschoengastia entomyza Womersley 1952

Neoschoengastia gallinarum ( Hatori, 1920)

Neoschoengastia hexasternosetosa (Chen and Hsu, 1958)

Neoschoengastia longipes Nadchatram, 1967

Neoschoengastia longitarsalis Schluger and Belskaya, 1966

Neoschoengastia posekanyi Wharton and Hardcastle, 1946

= Schoengastia (Ascoschoengastia) rectangulare Womersley 1952

= Neoschoengastia struthidia Womersley 1952

Neoschoengastia salmi ( Oudemans, 1922)

Neoschoengastia solitus Nadchatram, 1967

Neoschoengastia solomonis Wharton and Hardcastle, 1946

Neoschoengastia stekolnikovi ( Kalúz, 2016) , comb. nov.

Neoschoengastia thomasi (Radford, 1946)

Neoschoengastia vietnamensis Kalúz, 2016

Genus Ornithogastia Vercammen-Grandjean, 1960

Ornithogastia riversi ( Wharton and Hardcastle, 1946)

Genus Parascoschoengastia Vercammen-Grandjean, 1960

Parascoschoengastia heynemani (Nadchatram and Upham, 1966)

Parascoschoengastia monticola ( Wharton and Hardcastle, 1946)

Genus Riedlinia Oudemans, 1914

Riedlinia lipoxena ( Womersley, 1952)

Genus Schoengastia Oudemans, 1910

Schoengastia baguioensis Brown and Goff, 1988

Schoengastia crossi Nadchatram and Wooster, 1985

Schoengastia huxsolli Nadchatram, Goff and Thanalukshumi, 1980

Schoengastia kanhaensis Mitchell and Nadchatram, 1966

Schoengastia obtusispura Wang, 1962

Schoengastia palmata Domrow, 1962

Schoengastia propria Audy and Womersley, 1957

Schoengastia pseudoschuffneri ( Walch, 1927)

Schoengastia psorakari Nadchatram and Gentry, 1964

Schoengastia schuffneri ( Walch, 1922)

= Schoengastia katonis Womersley and Heaslip, 1943

= Schoengastia pusilla Womersley, 1944

Schoengastia sulawesiensis Goff, Durden and Whitaker, 1986

Schoengastia vandersandei ( Oudemans, 1905)

= Schoengastia blestowei Gunther, 1939

= Schoengastia blestowei var. megapodius Womersley and Heaslip, 1943

Schoengastia vieta Gater, 1932

= Schoengastia maldiviensis Radford, 1946

Genus Schoutedenichia Jadin and Vercammen-Grandjean, 1954

Subgenus Schoutedenichia Jadin and Vercammen-Grandjean, 1954

Schoutedenichia (Schoutedenichia) bisetosa Domrow, 1962

Schoutedenichia (Schoutedenichia) centralkwangtunga (Mo, Chen, Ho and Li, 1959)

= Schoutedenichia alongensis Schluger et al., 1960

Schoutedenichia (Schoutedenichia) trisetosa Upham and Nadchatram, 1968

Schoutedenichia (Schoutedenichia) vercammeni Audy, 1956

Genus Susa Audy and Nadchatram, 1960

Susa chiropteraphilus (Brown, 1997) , comb. nov.

Susa debilis ( Gater, 1932)

Susa labuanensis ( Womersley, 1952)

Susa macdonaldi Audy and Nadchatram 1960

Susa masawanensis (Brown, 1998) , comb. nov.

Susa palawanensis (Brown and Goff, 1988) , comb. nov.

Susa prachongae ( Lakshana, 1969)

Susa reidi Audy and Nadchatram 1960

Susa traubi Nadchatram and Lakshana, 1965

Genus Trisetica Traub and Evans, 1950

Trisetica melvini Traub and Evans, 1950

Genus Trombigastia Vercammen-Grandjean and Brennan, 1957

Trombigastia bishopi (Vercammen-Grandjean, 1964)

Trombigastia chrotogalis Chau, 2005

Trombigastia harrisoni ( Womersley, 1952)

Trombigastia rousetti Vercammen-Grandjean and Fain, 1958

Trombigastia tehminae Brown, 1997

Genus Walchiella Fuller, 1952

Walchiella beauforti Domrow, 1963

Walchiella bishopae Brown, 1998

Walchiella calunosa Traub and Audy, 1954

Walchiella hanseni Nadchatram and Lakshana, 1965

Walchiella harinastai Suzuki, 1980

Walchiella impar ( Gunther, 1939)

= Euschoengastia asonluca Traub and Audy, 1954

Walchiella lacunosa ( Gater, 1932)

Walchiella lobiensis Brown and Goff, 1988

Walchiella nadchatrami ( Womersley, 1952)

Walchiella oudemansi ( Walch, 1922)

= Neoschoengastia bodensis Gunther, 1940

Walchiella sarawakensis ( Womersley, 1952)

Walchiella traubi ( Womersley, 1952)

Tribe Trombiculini Vercammen-Grandjean, 1960

Genus Blankaartia Oudemans, 1911

Subgenus Blankaartia Oudemans, 1911

Blankaartia (Blankaartia) acuscutellaris ( Walch, 1922)

Subgenus Megaciella Vercammen-Grandjean, 1960

Blankaartia (Megaciella) kwanacara ( Chen and Hsu, 1957)

Genus Chiroptella Vercammen-Grandjean, 1960

Chiroptella audyi ( Vercammen-Grandjean and Nadchatram, 1965)

Chiroptella giga ( Schluger et al., 1959)

Chiroptella insolli ( Philip and Traub, 1950)

Chiroptella niehoffi (Domrow, 1962)

Chiroptella nocticola Nadchatram, 1966

Chiroptella nudusetosa Brown, 1998

Chiroptella piercei ( Ewing, 1931)

Chiroptella revelae ( Audy, 1952)

Chiroptella sandoshami Nadchatram, 1966

Chiroptella heidemani Brown, 1997

Genus Ericotrombidium Vercammen-Grandjean, 1966

Ericotrombidium cosmetopode (Vercammen-Grandjean and Langston, 1971) , comb. nov.

Genus Eutrombicula Ewing, 1938

Eutrombicula booliati (Audy, 1956)

Eutrombicula gigarara (Brown, 1997) , comb. nov.

Eutrombicula hirsti ( Sambon, 1927)

= Trombicula hakei Radford, 1946

Eutrombicula parmifera ( Womersley, 1952)

Eutrombicula rara ( Walch, 1924)

Eutrombicula scincoides ( Womersley, 1944)

Eutrombicula subrara (Audy, 1956)

Eutrombicula wichmanni ( Oudemans, 1905)

= Trombicula pseudoakamushi Hatori, 1920

= Trombicula mediocris Kawamura and Yamaguchi, 1921

= Trombicula hatorii Womersley and Heaslip, 1943

= Trombicula (Eutrombicula) wichmanni var. tinghuensis Liang, Lo and Hwang, 1957

Genus Fonsecia Radford, 1942

Fonsecia celesteae Audy, 1956

Fonsecia dispaseta Nadchatram and Upham, 1966

Fonsecia fasciata Chau, 2005

Genus Grandjeana Koçak and Kemal, 2009

Grandjeana asiatica ( Wen and Corpuz-Raros, 1997)

Grandjeana calva (Domrow, 1962)

Grandjeana koongi (Domrow, 1962)

Grandjeana manjuyodensis (Brown, 1997)

Grandjeana taphozous ( Womersley, 1952)

Genus Heaslipia Ewing, 1944

Heaslipia gateri ( Womersley and Heaslip, 1943)

Genus Laotrombicula Stekolnikov, 2014

Laotrombicula fangumi Stekolnikov, 2014

Laotrombicula khunboromi Stekolnikov, 2014

Genus Leptotrombidium Nagayo, Miyagawa, Mitamura and Imamura, 1916

Leptotrombidium abramovi Stekolnikov, 2013

Leptotrombidium aenigmami Stekolnikov, 2013

Leptotrombidium akamushi ( Brumpt, 1910)

= Microtrombidium brumpti Hirst, 1915

Leptotrombidium allosetum Wang, Liao and Lin, 1981

Leptotrombidium alopeciatum Traub and Nadchatram, 1967

Leptotrombidium andrei Tanskul and Gingrich, 1986

Leptotrombidium arenicola Traub, 1960

Leptotrombidium arvinum (Schluger, 1960)

Leptotrombidium baluense (Traub and Audy, 1954)

Leptotrombidium binbium Traub and Lakshana, 1966

Leptotrombidium bochkovi Stekolnikov, 2013

Leptotrombidium bodense ( Gunther, 1940)

Leptotrombidium brinchangense Nadchatram and Upham, 1966

Leptotrombidium burmense ( Ewing, 1945)

Leptotrombidium clarum Vercammen-Grandjean and Langston, 1976

Leptotrombidium deliense ( Walch, 1922)

= Trombicula walchi Womersley and Heaslip, 1943

= Leptotrombidium (Leptotrombidium) deliense sinense Wen and Chen, 1984

= Leptotrombidium (Leptotrombidium) deliense microsetosa Zhao, Tang and Mo, 1986

Leptotrombidium dendrium Tanskul, 1991

Leptotrombidium elisbergi Traub and Lakshana, 1966

Leptotrombidium fletcheri ( Womersley and Heaslip, 1943)

= Leptotrombidium (Leptotrombidium) fletcheri francolini Wen and Xiang, 1984

Leptotrombidium fulleri ( Ewing, 1945)

= Trombicula (Leptotrombidium) pumilis Traub, Morrow and Lipovsky, 1958

Leptotrombidium fulmentum Vercammen-Grandjean and Langston, 1976

Leptotrombidium furcagaleala Vercammen-Grandjean and Langston, 1971

Leptotrombidium gentryi Nadchatram and Upham, 1966

Leptotrombidium globosum (Schluger, 1960)

Leptotrombidium gracipalpe (Schluger, 1960)

Leptotrombidium hanseni Traub and Lakshana, 1966

Leptotrombidium harrisoni Tanskul and Gingrich, 1986

Leptotrombidium horridum (Schluger, 1960)

Leptotrombidium imphalum Vercammen-Grandjean and Langston, 1976

= Leptotrombidium (Leptotrombidium) imphalum ceylonicum Vercammen-Grandjean and Langston, 1976

= Leptotrombidium (Leptotrombidium) imphalum sabahense Vercammen-Grandjean and Langston, 1976

= Leptotrombidium (Leptotrombidium) chiangraiensis Tanskul and Linthicum, 1997

Leptotrombidium insolitum Nadchatram and Upham, 1966

Leptotrombidium javani Vercammen-Grandjean and Langston, 1976

Leptotrombidium keukenschrijveri ( Walch, 1924)

Leptotrombidium kinabalui Stekolnikov, 2013

Leptotrombidium kundini Nadchatram and Upham, 1966

Leptotrombidium kunshui Wen and Xiang, 1984

Leptotrombidium labuani Vercammen-Grandjean and Langston, 1976

Leptotrombidium lanceolatum ( Womersley, 1952)

Leptotrombidium langati ( Audy and Womersley, 1957)

Leptotrombidium laoense Stekolnikov, 2013

Leptotrombidium linjeromae Vercammen-Grandjean and Langston, 1976

Leptotrombidium longimedian Brown, 1992

= Leptotrombidium (Leptotrombidium) mindanensis Brown, 1992

Leptotrombidium longisetum ( Womersley, 1952)

Leptotrombidium luzonense Vercammen-Grandjean and Langston, 1976

Leptotrombidium macacaphilum Brown, 1992

Leptotrombidium macacum ( Womersley, 1952)

= Mehracula roonwali Sinha, 1954

Leptotrombidium madiense Traub and Nadchatram, 1967

Leptotrombidium magnum (Schluger, 1960)

Leptotrombidium malayanum Nadchatram and Upham, 1966

Leptotrombidium megabodense Stekolnikov, 2013

Leptotrombidium megalangati Stekolnikov, 2013

Leptotrombidium megaloti Stekolnikov, 2013

Leptotrombidium miculum (Traub and Audy, 1954)

Leptotrombidium minului Stekolnikov, 2013

Leptotrombidium monstrosum (Schluger, 1960)

Leptotrombidium muridium ( Womersley, 1952)

Leptotrombidium namru Hadi, 1991

Leptotrombidium nguyenvanaii Phuong, 1972

Leptotrombidium oculascutum Brown, 1992

Leptotrombidium paradux Vercammen-Grandjean and Langston, 1976

= Leptotrombidium montanum Stekolnikov, 2004

Leptotrombidium paulum Vercammen-Grandjean and Langston, 1976

Leptotrombidium peniculatum Traub and Lakshana, 1966

Leptotrombidium periosum Traub and Nadchatram, 1967

Leptotrombidium phangi Upham and Nadchatram, 1968

Leptotrombidium philippinense Kolebinova, 1980

Leptotrombidium pilaltum (Traub and Audy, 1954)

Leptotrombidium pilosum Traub and Lakshana, 1966

Leptotrombidium pipellae (Traub and Audy, 1954)

Leptotrombidium rapmundi Nadchatram and Upham, 1966

Leptotrombidium rectanguloscutum ( Hsu and Chen, 1964)

Leptotrombidium saigoni Stekolnikov, 2013

Leptotrombidium sandfordi Goff, 1981

Leptotrombidium scanloni Traub and Lakshana, 1966

Leptotrombidium scutellare ( Nagayo, Miyagawa, Mitamura, Tamiya and Tenjin, 1921)

= Trombicula (Leptotrombidium) scutellaris var. basoglabrosis Chen, Hsu and Wang, 1956

Leptotrombidium serum Vercammen-Grandjean and Langston, 1976

Leptotrombidium sialkotense Vercammen-Grandjean and Langston, 1976

= Leptotrombidium (Leptotrombidium) jishoum Wen, Li, Zhang and Liao, 1988

Leptotrombidium spicapilum Vercammen-Grandjean and Langston, 1976

Leptotrombidium spicatum Traub, 1960

Leptotrombidium subangi Vercammen-Grandjean and Langston, 1976

Leptotrombidium subangulare Wen and Xiang, 1984

Leptotrombidium subobscurum Kolebinova, 1980

Leptotrombidium sylvestre ( Audy and Traub, 1950)

Leptotrombidium tardum Vercammen-Grandjean and Langston, 1976

Leptotrombidium tarsale (Traub and Audy, 1954)

Leptotrombidium taiyuanense Tian and Wen, 1984

Leptotrombidium tenompaki Stekolnikov, 2013

Leptotrombidium tenuipalpe Vercammen-Grandjean and Langston, 1976

Leptotrombidium tikhonovi Stekolnikov, 2013

Leptotrombidium tupaianum Vercammen-Grandjean and Langston, 1976

Leptotrombidium turdicola Vercammen-Grandjean and Langston, 1976

= Leptotrombidium (Leptotrombidium) muntiaci Xiang and Wen, 1984

= Leptotrombidium (Leptotrombidium) suense Wen, 1984

Leptotrombidium ului Stekolnikov, 2013

Leptotrombidium umbricola Nadchatram and Dohany, 1980

Leptotrombidium urogale Brown, 1992

Leptotrombidium vivericola Vercammen-Grandjean and Langston, 1976

Leptotrombidium waltoni Vercammen-Grandjean and Langston, 1976

Leptotrombidium yunlingense Yu, Yang, Zhang, Zi and Hu, 1981

Genus Lorillatum Nadchatram, 1963

Lorillatum attapinum ( Schluger et al., 1963)

Lorillatum flagelliferum (Traub and Audy, 1954)

Lorillatum hekouensis Yu, Chen and Lin, 1996

Lorillatum kianjoei (Nadchatram and Traub, 1964)

Lorillatum mastigophorum (Nadchatram, 1963)

Lorillatum oreophilum (Nadchatram and Traub, 1964)

Lorillatum panitae (Nadchatram and Traub, 1964)

Lorillatum tuhanum (Traub and Audy, 1954)

Genus Microtrombicula Ewing, 1950

Microtrombicula audyi Vercammen-Grandjean, 1965

Microtrombicula batui ( Philip and Traub, 1950)

Microtrombicula chamlongi Nadchatram and Kethley, 1974

Microtrombicula dasia Brown, 1999

Microtrombicula eltoni (Audy, 1956)

Microtrombicula fieldi (Audy, 1956)

Microtrombicula fulgida ( Schluger et al., 1963)

Microtrombicula lepidodactilus Brown, 1999

Microtrombicula megaderma Brown, 1997

Microtrombicula munda ( Gater, 1932)

Microtrombicula nadchatrami Vercammen-Grandjean, 1965

Microtrombicula octaintercoxalaea Brown, 1997

Microtrombicula spelaea Goff, 1990

Microtrombicula spicea ( Gater, 1932)

Microtrombicula tweediei (Audy, 1956)

Microtrombicula vitosa ( Schluger et al., 1963)

Genus Miyatrombicula Sasa, Kawashima and Egashira, 1952

Miyatrombicula benensoni Tanskul and Nadchatram, 1983

Genus Myotrombicula Womersley and Heaslip, 1943

Myotrombicula armigera Brown, 1997

Myotrombicula medwayi Nadchatram and Lakshumy, 1969

Myotrombicula oconnori Brown, 1997

Myotrombicula philipi ( Womersley, 1952)

Myotrombicula vercammeni Nadchatram and Lakshana, 1965

Myotrombicula womersleyi Vercammen-Grandjean, 1968

Genus Neotrombicula Hirst, 1925

Neotrombicula anax Audy and Womersley, 1957

Neotrombicula elegans Schluger, 1966

Neotrombicula fujigmo ( Philip and Fuller, 1950)

Neotrombicula scorpionis Lakshana, 1966

Neotrombicula weni ( Wang, 1964)

Genus Novotrombicula Womersley and Kohls, 1947

Novotrombicula owiensis Womersley and Kohls, 1947

Genus Octasternala Brown, 1990

Octasternala taphozousa Brown, 1990

Genus Rudnicula Vercammen-Grandjean, 1964

Rudnicula becki Nadchatram, 1968

Rudnicula dimolinae ( Audy, 1952)

Rudnicula hexasternalaea Brown, 1997

Rudnicula leytensis Brown, Goff and Nadchatram, 1988

Genus Sasatrombicula Vercammen-Grandjean, 1960

Sasatrombicula keechongi ( Nadchatram and Mitchell, 1965)

Sasatrombicula leegoffi Brown, 1997

Sasatrombicula siamensis ( Nadchatram and Mitchell, 1965)

Genus Toritrombicula Sasa, Hayashi and Kawashima, 1953

Toritrombicula asa Nadchatram, 1967

Toritrombicula densipiliata ( Walch, 1922)

Toritrombicula dicrura Chau, 2005

Toritrombicula dumbletoni ( Womersley and Audy, 1957)

Toritrombicula robusta ( Gunther, 1941)

Toritrombicula samara (Radford, 1953)

Toritrombicula uphami Nadchatram, 1967

Toritrombicula vorca (Traub and Audy, 1954)

Genus Trombicula Berlese, 1905

Subgenus Cotrombicula Vercammen-Grandjean, 1960

Trombicula (Cotrombicula) gressitti Nadchatram and Wilson, 1965

Trombicula (Cotrombicula) macclurei Vercammen-Grandjean and Nadchatram, 1963

Trombicula (Cotrombicula) megaderma Vercammen-Grandjean and Nadchatram, 1963

Subgenus Trombicula Berlese, 1905

Trombicula (Trombicula) minor Berlese, 1905

Genus Trombiculindus Radford, 1948

Trombiculindus alethrix (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus armatum ( Traub, Nadchatram and Lakshana, 1968)

Trombiculindus atractimorphe ( Traub, Nadchatram and Lakshana 1968)

Trombiculindus cuneatum Traub and Evans, 1951

Trombiculindus cuteanum ( Vercammen-Grandjean and Langston, 1976) , comb. nov.

Trombiculindus foliaceum Traub and Evans, 1951

Trombiculindus frondosum (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus gateri (Traub and Nadchatram, 1967)

Trombiculindus hastatum ( Gater, 1932) , comb. nov.

Trombiculindus griselda ( Schluger et al., 1963)

Trombiculindus imbricatum (Traub and Nadchatram, 1967)

Trombiculindus kosapani Stekolnikov, 2016

Trombiculindus lepismatum (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus limi (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus macrosphenum ( Traub, Nadchatram and Lakshana, 1968)

Trombiculindus manooni ( Suzuki, 1980)

Trombiculindus maxwelli (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus paniculatum ( Traub, Nadchatram and Lakshana, 1968)

Trombiculindus roseannleilaniae ( Brown, 1992) , comb. nov.

Trombiculindus santasirii ( Suzuki, 1980)

Trombiculindus sarisatum (Traub and Nadchatram, 1967) , comb. nov.

Trombiculindus sibynatum ( Traub, Nadchatram and Lakshana, 1968)

Trombiculindus thurmani ( Traub, Nadchatram and Lakshana, 1968)

Trombiculindus vanpeeneni ( Hadi and Carney, 1977) , comb. nov.

Trombiculindus variaculum (Traub and Nadchatram, 1967)

Trombiculindus yooni (Traub and Nadchatram, 1967) , comb. nov.

Genus Vatacarus Southcott, 1957

Vatacarus ipoides Southcott, 1957

Genus Vercammenia Audy and Nadchatram, 1957

Vercammenia hendricksoni Audy and Nadchatram, 1957

Incertae sedis

Eutrombicula gymnodactyla Womersley and Kohls, 1947

Trombicula mediocris Berlese, 1912

Abonnenc, E. & Taufflieb, R. (1957) Deux trombiculides nouveaux de la region du Tchad, Euschoengastia pilosa n. sp. et Trombicula youhensis n. sp., presence de Schoengastia radfordi Jadin et Vercammen-Grandjean, 1954. Bulletin de la Societe de pathologie exotique, 50, 556 - 567.

Andre, M. (1954) Description d'une nouvelle larve d' Ascoschongastia (Acarien) parasite de rats en Indochine. Bulletin du Museum National d'Histoire Naturelle, Paris, 2 e Series, 26, 200 - 203.

Audy, J. R. & Traub, R. (1950) Trombicula sylvestris sp. nov. (Acarina, Trombiculidae) from Malaya. Bulletin of the Raffles Museum, 23, 325 - 327.

Audy, J. R. (1952) Trombiculid mites infesting bats in Malaya, with descriptions of three new species. Bulletin of the Raffles Museum, 24, 132 - 159.

Audy, J. R. (1954) Malaysian Parasites-IX. Notes on the taxonomy of trombiculid mites, with description of a new subgenus. Studies from the Institute for Medical Research, Federation of Malaya, 26, 123 - 170. https: // doi. org / 10.5962 / bhl. title. 119064

Audy, J. R. & Nadchatram, M. (1960) Malaysian Parasites-XXXIX. Susa, new genus related to Ascoschongastia Ewing (Acarina, Trombiculidae), with descriptions of two new species. Studies from the Institute for Medical Research, Federation of Malaya, 29, 154 - 162.

Berlese, A. (1905) Acari nuovi. Manipulus IV (Acari de Giava). Redia, 2, 154 - 176.

Berlese, A. (1912) Trombidiidae. Prospetto dei generi edelle specie finora noti. Redia, 8, 1 - 291.

Brown, W. A., Goff, M. L. & Nadchatram, M. (1988) The bat-infesting genus Rudnicula (Acari: Trombiculidae), with description of a new species from the Philippine Islands and a key to the species. Journal of Medical Entomology, 25, 17 - 19. https: // doi. org / 10.1093 / jmedent / 25.1.17

Brown, W. A. (1990) A new genus and species of chigger (Acari: Trombiculidae) from the Philippine Islands. International Journal of Acarology, 16, 23 - 25. https: // doi. org / 10.1080 / 01647959008683860

Brown, W. A. (1992) Description of six new species of the genus Leptotrombidium from Mindanao Island and notes on the medically important genera of chiggers (Acari: Trombiculidae) of the Philippine Islands. Journal of Medical Entomology, 29, 284 - 292. https: // doi. org / 10.1093 / jmedent / 29.2.284

Brown, W. A. (1999) Two new species of Microtrombicula (Acari: Trombiculidae) from the Philippines. International Journal of Acarology, 25, 19 - 22. https: // doi. org / 10.1080 / 01647959908683607

Brumpt, E. (1910) Precis de parasitologie. 2 nd Edition. Masson et Cie, Paris, 915 pp.

Chau, N. V. (2005) Three new chigger species (Acariformes: Trombiculidae) collected on mammal, bird and reptile from Vietnam. Tap chi Sinh hoc, 27 (2), 8 - 15. [in Vietnamese] https: // doi. org / 10.15625 / 0866 - 7160 / v 27 n 2.5254

Chen, H. T., Hsu, P. K. & Wang, T. (1956) Five species and varieties of trombiculid mites from China. Acta Zoologica Sinica, 8, 149 - 160. [in both Chinese and English]

Chen, H. T. & Hsu, P. K. (1957) Scrub mites from mammals and birds including the description of three new species. Acta Zoologica Sinica, 9, 379 - 404. [in Chinese]

Chen, H. T. & Hsu, P. K. (1965) Seven new species of trombiculid mites belonging to the genera Doloisia and Paradoloisia (Acari, Trombiculidae). Acta Zoologica Sinica, 17, 283 - 297. [in both Chinese and English]

Domrow, R. & Nadchatram, M. (1962) Three new species of Doloisia from S. E. Asia (Acarina, Trombiculidae). Acarologia, 4, 577 - 585.

Domrow, R. (1963) The genus Walchiella (Acarina, Trombiculidae). Proceedings of the Linnean Society of New South Wales, 87, 105 - 115.

Ewing, H. E. (1931) A catalogue of the Trombiculinae, or chigger mites, of the New World with new genera and species and a key to the genera. Proceedings of the United States National Museum, 80, 1 - 19. https: // doi. org / 10.5479 / si. 00963801.80 - 2908.1

Ewing, H. E. (1945) Two new trombiculid mite larvae (chiggers) from Burma. Proceedings of the Entomological Society of Washington, 47, 63 - 65.

Fuller, H. S. (1952) The mite-larvae of the family Trombiculidae in the Oudemans collection: taxonomy and medical importance. Zoologische Verhandelingen, 18, 1 - 261.

Gater, B. A. R. (1932) Malayan trombidiid larvae, part I. (Acarina: Trombidiidae) with descriptions of seventeen new species. Parasitology, 24, 143 - 174. https: // doi. org / 10.1017 / S 0031182000020515

Goff, M. L., Durden, L. A. & Whitaker, J. O. (1986) A new species of Schoengastia (Acari: Trombiculidae) from mammals collected in Sulawesi, Indonesia. International Journal of Acarology, 12, 91 - 93. https: // doi. org / 10.1080 / 01647958608683448

Goff, M. L. (1990) A new species of Microtrombicula (Acari: Trombiculidae) from bats (Chiroptera: Pteropidae) collected in Indonesia. International Journal of Acarology, 16, 175 - 177. https: // doi. org / 10.1080 / 01647959008683528

Gunther, C. E. M. (1939) Trombidiid larvae in New Guinea (Acarina: Trombidiidae). Proceedings of the Linnean Society of New South Wales, 64, 73 - 96.

Gunther, C. E. M. (1940) Four larval Trombidiidae from British North Borneo (Acarina: Trombidiidae). Proceedings of the Linnean Society of New South Wales, 65, 479 - 483.

Gunther, C. E. M. (1941) Two new trombidiid larvae from New Guinea (Acarina: Trombidiidae). Proceedings of the Linnean Society of New South Wales, 66, 157 - 159.

Hadi, T. R. & Carney, W. P. (1977) Two new trombiculid mites from mammals of South Vietnam, with locality records for six additional species (Acarina: Trombiculidae). Journal of Medical Entomology, 14, 455 - 460. https: // doi. org / 10.1093 / jmedent / 14.4.455

Hadi, T. (1991) Chiggers (Acarina, Trombiculidae) of Maxomys bartelsii found in West Java, Indonesia, with descriptions of two new species. Treubia, 30, 95 - 101.

Hatori, J. (1920) Report on the tsutsugamushi disease in Formosa. Taiwan Igakki Zasshi, 209, 317 - 352.

Hirst, S. (1921) On three new parasitic mites (Leptus, Schongastia and Demodex). Annals and Magazine of Natural History, Series 9, 7, 37 - 39. https: // doi. org / 10.1080 / 00222932108632487

Hirst, S. (1925) On a harvest bug (Leeuwenhoekia australiensis, sp. n.) attacking human beings at Sidney, New South Wales. Transactions of the Royal Society of Tropical Medicine and Hygien e, 19, 150 - 152. https: // doi. org / 10.1016 / S 0035 - 9203 (26) 80005 - 7

Hsu, P. K. & Chen, H. T. (1964) Eight new species of scrub mites from Kwangtung. Acta Entomologica Sinica, 13, 737 - 752. [in both Chinese and English]

Kaluz, S. K., Hung, N. M., Capek, M. & Literak, I. (2016) Two new species and new records of chiggers (Acari: Leeuwenhoeki- idae, Trombiculidae) from birds in Vietnam. Zootaxa, 4061 (5), 483 - 503. https: // doi. org / 10.11646 / zootaxa. 4061.5.2

Kawamura, R. & Yamaguchi, M. (1921) Ueber die Tsutsugamushi-Krankheit in Formosa, zugleich eine vergleichende Studie derselben mit der in Nordjapan. Kitasato archives of experimental medicine, 4, 169 - 206.

Kocak, A. O. & Kemal, M. (2009) A replacement name in the family Trombiculidae in Acarina. Centre for Entomological Studies Ankara, Miscellaneous Papers, 147 - 148, 15 - 16.

Kolebinova, M. G. (1980) Three new species from genus Leptotrombidium (Acarina, Trombiculidae). Acta Zoologica Bulgarica, 16, 69 - 75.

Lakshana, P. (1966) A new species of trombiculid mite infesting scorpions in Thailand (Acarina: Trombiculidae). Journal of Medical Entomology, 3, 258 - 260. https: // doi. org / 10.1093 / jmedent / 3.3 - 4.258

Lakshana, P. (1969) Susa Audy & Nadchatram, a subgenus of Cheladonta Lipovsky et al. (new status), with descriptions of two new species of Cheladonta from Thailand. Journal of Medical Entomology, 6, 135 - 139. https: // doi. org / 10.1093 / jmedent / 6.2.135

Liang, B. L., Lo, T. H. & Hwang, Y. Y. (1957) A new species and a new variety of trombiculid mites (Acarina: Trombiculidae) from Kwangtung province. Acta Zoologica Sinica, 9, 221 - 226. [in both Chinese and English]

Mitchell, C. J. & Nadchatram, M. (1966) Seven new species of chiggers from Central India, with a redescription of Leptotrombidium (L.) pelta (Womersley, 1952) (Acarina: Trombiculidae). Journal of Medical Entomology, 3, 61 - 77. https: // doi. org / 10.1093 / jmedent / 3.1.61

Nadchatram, M. & Domrow, R. (1964) The intranasal species of Laurentella (Acarina: Trombiculidae). Journal of Medical Entomology, 1, 29 - 39. https: // doi. org / 10.1093 / jmedent / 1.1.29

Nadchatram, M. & Gentry, J. W. (1964) A new species of scrub-itch mite from Malaya, with notes on ecology (Acarina, Trombiculidae). Journal of Medical Entomology, 1, 1 - 4. https: // doi. org / 10.1093 / jmedent / 1.1.1

Nadchatram, M. & Mitchell, C. J. (1965) New bat chiggers from Thailand and the Solomon Islands with notes on the subgenus Sasatrombicula Vercammen-Grandjean (Acarina: Trombiculidae). Journal of Medical Entomology, 2, 70 - 74. https: // doi. org / 10.1093 / jmedent / 2.1.70

Nadchatram, M. & Wilson, N. (1965) Four new species of bat chiggers from New Guinea (Acarina: Trombiculidae). Journal of Medical Entomology, 2, 217 - 224. https: // doi. org / 10.1093 / jmedent / 2.3.217

Nadchatram, M. (1968) Notes on the genus Rudnicula (Acarina, Trombiculidae) with description of a new species from Malaysia. Acarologia, 10, 467 - 471.

Nadchatram, M. & Lakshumy, G. T. (1969) A new species and new record of chiggers from West Malaysia (Prostigmata: Trombiculidae). Journal of Medical Entomology, 6, 283 - 285. https: // doi. org / 10.1093 / jmedent / 6.3.283

Nadchatram, M. & Traub, R. (1971) Chiggers of the genus Helenicula of the Old World including descriptions of 9 new species (Acarina: Prostigmata, Trombiculidae). Journal of Medical Entomology, 8, 562 - 597. https: // doi. org / 10.1093 / jmedent / 8.5.562

Nadchatram, M. & Kethley, J. (1974) A collection of reptilian chiggers from Thailand with descriptions of three new species (Acari: Prostigmata, Trombiculidae) and preliminary notes on their biologies. Journal of Medical Entomology, 11, 581 - 587.

Nadchatram, M., Goff, M. L. & Thanalukshumi, P. (1980) The genus Schoengastia (Acari: Trombiculidae) in the Asiatic-Pacific region. Journal of Medical Entomology, 17, 268 - 281. https: // doi. org / 10.1093 / jmedent / 17.3.268

Nadchatram, M. & Dohany, A. L. (1980) Leptotrombidium (Leptotrombidium) umbricola, new species, a probable vector of scrub typhus in Peninsular Malaysia. Japanese Journal of Medical Science and Biology, 33, 277 - 282. https: // doi. org / 10.7883 / yoken 1952.33.277

Nadchatram, M. & Wooster, M. T. (1985) Schoengastia crossi, a new species of scrub itch chigger from the Philippines (Acari, Prostigmata, Trombiculidae). Tropical Biomedicine, 2, 59 - 62.

Nagayo, M., Miyagawa, Y., Mitamura, T., Tamiya, T. & Tenjin, S. (1921) Five species of tsutsugamushi (the carrier of Japanese river fever) and their relation to the tsutsugamushi disease. American Journal of Hygiene, 1, 569 - 591. https: // doi. org / 10.1093 / oxfordjournals. aje. a 118053

Oudemans, A. C. (1905) Acarologische Aanteekeningen. XVI. Entomologische Berichten, Amsterdam, 1, 216 - 218. https: // doi. org / 10.5962 / bhl. part. 1124

Oudemans, A. C. (1910) Acarologische Aanteekeningen. XXXIII. Entomologische Berichten, Amsterdam, 3, 83 - 90. https: // doi. org / 10.5962 / bhl. part. 1133

Oudemans, A. C. (1922) Acarologische Aanteekeningen. LXVII. Entomologische Berichten, Amsterdam, 6, 81 - 84.

Philip, C. B. (1947) Ascoschoengastia bushlandi, a new species of chigger mite from Dutch New Guinea. Journal of Parasitology, 33, 387 - 390. https: // doi. org / 10.2307 / 3273672

Philip, C. B. & Traub, R. (1950) Two new species of trombiculid mites from Malayan bats. Journal of Parasitology, 36, 29 - 33. https: // doi. org / 10.2307 / 3273487

Philip, C. B. & Fuller, H. S. (1950) The harvest mites (' Akidani') of Japan and the Far East and their relationship to the Autumnalis group of trombiculid mites. Parasitology, 40, 50 - 57. https: // doi. org / 10.1017 / S 0031182000017856

Phuong, T. T. M. (1972) Description d'une nouvelle espece de Leptotrombidium (L.) du Sud Viet-Nam. Bulletin de la Societe de pathologie exotique, 6, 889 - 893.

Radford, C. D. (1942) The larval Trombiculinae (Acarina: Trombidiidae) with descriptions of twelve new species. Parasitology, 34, 55 - 81. https: // doi. org / 10.1017 / S 0031182000015997

Sambon, L. W. (1927) XIX. - The scrub itch-mite of north Queensland: a new species of Trombicula. Annals and Magazine of Natural History, Series 9, 20, 157 - 161. https: // doi. org / 10.1080 / 00222932708655577

Schluger, E. G., Grochovskaja, I. M., Ngu, D. V., Hoe, N. X. & Tung, D. K. (1959) New species of chiggers (Acariformes) from bats of North Viet-Nam. Zoologicheskiy Zhurnal, 38, 418 - 425. [in Russian]

Schluger, E. G., Grochovskaja, I. M., Ngu, D. V., Hoe, N. X. & Tung, D. K. (1961) The trombiculid mites (chigger mites) of the genera Doloisia Oudemans, 1910, and Traubacarus Audy et Nadchatram, 1957 (Acariformes, Trombiculidae) from North Viet-Nam. Entomologicheskoe Obozrenie, 40, 448 - 453. [in Russian]

Schluger, E. G., Grochovskaja, I. M., Ngu, D. V., Hoe, N. X. & Tung, D. K. (1963) Chigger-mites of the genus Trombicula (Acariformes, Trombiculidae) from Democratic Republic Viet Nam. Entomologicheskoe Obozrenie, 42, 691 - 701. [in Russian]

Schluger, E. G. & Belskaya, G. S. (1966) New species of chigger mites (Acariformes, Trombiculidae) from birds of Turkmenia. Izvestiia Akademii Nauk Turkmenskoj SSR, Seria biologicheskikh nauk, 1, 64 - 70. [in Russian]

Sinha, T. B. (1954) New species of larval mites from India (Acarina: Trombiculidae). Parasitology, 44, 329 - 337. https: // doi. org / 10.1017 / s 0031182000018977

Southcott, R. V. (1957) On Vatacarus ipoides n. gen., n. sp. (Acarina: Trombidioidea). A new respiratory endoparasite from a Pacific sea-snake. Transactions of the Royal Society of South Australia, 80, 165 - 176.

Stekolnikov, A. A. (2004) Variability in Leptotrombidium europaeum and two new related chigger mite species (Acari: Trombiculidae) from Caucasus. Parazitologiya, 38, 388 - 405. [in Russian]

Stekolnikov, A. A. (2013) Leptotrombidium (Acari: Trombiculidae) of the World. Zootaxa, 3728 (1), 1 - 173. https: // doi. org / 10.11646 / zootaxa. 3728.1.1

Stekolnikov, A. A. (2014) A new genus and two new species of chigger mites (Acari: Trombiculidae) from the Laotian rock-rat Laonastes aenigmamus Jenkins, Kilpatrick, Robinson & Timmins (Rodentia: Diatomyidae). Systematic Parasitology, 87, 21 - 31. https: // doi. org / 10.1007 / s 11230 - 013 - 9453 - 4

Suzuki, H. (1980) Studies on the parasite fauna of Thailand. 4. Five new species of trombiculid mites found in Thailand (Prostigmata: Trombiculidae). Tropical Medicine, 22, 75 - 87.

Takekawa, S. (1945) On a new species of Trombicula, T. soekaboemiensis, n. sp., found in Java. Bulletin Nampogun Boekikyusui-Bu, Singapore, 126, 1 - 5.

Tanskul, P. & Nadchatram, M. (1983) Notes on the genus Miyatrombicula (Acari: Prostigmata: Trombiculidae), with description of a new species from Thailand. Journal of Medical Entomology, 20, 597 - 600. https: // doi. org / 10.1093 / jmedent / 20.6.597

Tanskul, P. & Gingrich, J. B. (1986) Two new species of Leptotrombidium (Acari: Actinedida: Trombiculidae), probable vectors of scrub typhus in Thailand. Journal of Medical Entomology, 23, 661 - 665. https: // doi. org / 10.1093 / jmedent / 23.6.661

Tanskul, P. (1991) A new species of Leptotrombidium (Acari: Trombiculidae) collected from black plates in Thailand. Journal of Medical Entomology, 28, 139 - 141. https: // doi. org / 10.1093 / jmedent / 28.1.139

Tanskul, P. & Linthicum, K. J. (1997) A new species of Leptotrombidium (Acari: Trombiculidae) collected in active rice fields in northern Thailand. Journal of Medical Entomology, 34, 368 - 371. https: // doi. org / 10.1093 / jmedent / 34.3.368

Tian, Q. & Wen, T. (1984) Two new species of the genus Leptotrombidium from Shanxi Province (Acariformes: Trombiculidae). In: Wen, T. H. (Ed.), Sand mites of China (Acariformes: Trombiculidae and Leeuwenhoekiidae). Hue Lin Publishing House, Shanghai, pp. 43 - 45. [in Chinese]

Traub, R. & Sundermeyer, E. (1950) A new species of Ascoschoengastia from north Burma (Acarina: Trombiculidae). Proceedings of the Helminthological Society of Washington, 17, 35 - 38.

Traub, R. & Evans, T. M. (1951) Two new species of Trombiculindus Radford from North Burma (Acarina: Trombiculidae). Annals of the Entomological Society of America, 44, 262 - 268. https: // doi. org / 10.1093 / aesa / 44.2.262

Traub, R., Morrow, M. L. & Lipovsky, L. J. (1958) New species of chiggers from Korea. Proceedings of the Entomological Society of Washington, 60, 145 - 166.

Traub, R. (1960) Malaysian Parasites-XLV. Two new species of chiggers of the genus Leptotrombidium (Acarina, Trombiculidae). Studies from the Institute for Medical Research, Federation of Malaya, 29, 198 - 204.

Traub, R. & Lakshana, P. (1966) Some chiggers of the subgenus Leptotrombidium from Thailand, with descritions of new species (Acarina, Trombiculidae). Journal of Medical Entomology, 3, 271 - 292. https: // doi. org / 10.1093 / jmedent / 3.3 - 4.271

Traub, R., Nadchatram, M. & Lakshana, P. (1968) New species of chiggers of the subgenus Trombiculindus from Thailand (Acarina, Trombiculidae- Leptotrombidium). Journal of Medical Entomology, 5, 363 - 374. https: // doi. org / 10.1093 / jmedent / 5.3.363

Upham, R. W. & Nadchatram, M. (1968) Three new species of trombiculid mites from Malaysia with a redescription and notes on Leptotrombidium muridia (Womersley) (Acarina, Trombiculidae). Journal of Medical Entomology, 5, 195 - 203. https: // doi. org / 10.1093 / jmedent / 5.2.195

Vercammen-Grandjean, P. H. & Brennan, J. M. (1957) Eight new chiggers from East Africa and a new genus, Trombigastia (Acarina: Trombiculidae). Annals of the Entomological Society of America, 50, 484 - 496. https: // doi. org / 10.1093 / aesa / 50.5.484

Vercammen-Grandjean, P. H. & Fain, A. (1958) Les Trombiculidae parasites de Chiropteres. Revision du genre Trombigastia, description d'un Myotrombicula. Annales de parasitologie humaine et comparee, 33, 5 - 35.

Vercammen-Grandjean, P. H. (1960) Introduction a un essai de classification rationelle des larves de Trombiculinae Ewing, 1944 (Acarina: Trombiculidae). Acarologia, 2, 469 - 471.

Vercammen-Grandjean, P. H. & Nadchatram, M. (1963) Le genre Trombicula et ses sous-genres (Trombiculidae - Acarina). Acarologia, 5, 384 - 393.

Vercammen-Grandjean, P. H. & Nadchatram, M. (1965) New considerations about the genus Riedlinia Oudemans, 1914. Reinstatement of the genus Trombigastia Vercammen-Grandjean and Brennan, 1957. Erection of a new genus, Bishoplinia (Acarina: Trombiculidae). Acarologia, 7 (suppl.), 317 - 324.

Vercammen-Grandjean, P. H. (1966) Revision of the genus Herpetacarus Vercammen-Grandjean, 1960 (Trombiculidae - Acarina). Acarologia, 8, 631 - 674.

Vercammen-Grandjean, P. H. (1967) Notes on the Trombiculidae. Acarologia, 9, 127 - 134.

Vercammen-Grandjean, P. H. & Langston, R. L. (1976) The chigger mites of the World (Acarina: Trombiculidae et Leeuwenhoekiidae). III. Leptotrombidium complex. George Williams Hooper Foundation, University of California, San Francisco, California, 1061 pp.

Walch, E. W. (1922) Over Trombicula deliensis n. sp., vermoedelijke overbrengster der Pseudotyphus, en andere Trombiculae van Deli (Eerste Mededeeling). Geneeskundig Tijdschrift voor Nederlandsch-Indie, 62, 530 - 588.

Walch, E. W. (1924) Over de Trombiculae, welke de pseudotyphus overbrengen, en na verwante mijten uit Deli (2 e mededeeling). Geneeskundig Tijdschrift voor Nederlandsch-Indie, 64, 499 - 529.

Walch, E. W. (1927) Nederlandsch-Indische Trombiculae en verwante mijten (derde mededeeling). Soorten uit de Lamprongsche Districten en de Omgeving van Makassar. Geneeskundig Tijdschrift voor Nederlandsch-Indie, 67, 922 - 933.

Wang, D. C. (1962) On three new species and one new record of the trombiculid mites from China. Acta Entomologica Sinica, 11, 269 - 276. [in Chinese]

Wang, D. C. (1964) Five new species and one new record of chiggers (Acarina: Trombiculidae). Acta Entomologica Sinica, 13, 88 - 100. [in Chinese]

Wang, D. Q., Liao, H. R. & Lin, Z. H. (1981) Five new trombiculid mites of the genus Leptotrombidium (Acarina: Trombiculidae). Acta Zootaxonomica Sinica, 6, 44 - 52. [in Chinese]

Wen, T. H. & Chen, J. (1984) A new subspecies of Leptotrombidium deliense from Penghu, Taiwan Province (Acariformes: Trombiculidae). In: Wen, T. H. (Ed.), Sand mites of China (Acariformes: Trombiculidae & Leeuwenhoekiidae). Hue Lin Publishing House, Shanghai, pp. 175 - 176. [in Chinese]

Wen, T. H., Li, G., Zhang, X. & Liao, H. (1988) A new species of sand mite from the western part of Hunan Province, China (Acariformes: Trombiculidae). Entomotaxonomia, 10, 107 - 113. [in Chinese]

Wen, T. H. & Corpuz-Raros, L. A. (1997) Notes on the sand mite genus Diplectria and description of new species from the Philippines (Acariformes: Trombiculidae). Philippine Entomologist, 11, 25 - 31.

Wharton, G. W. & Hardcastle, A. B. (1946) The genus Neoschoengastia (Acarinida: Trombiculidae) in the Western Pacific area. Journal of Parasitology, 32, 286 - 322. https: // doi. org / 10.2307 / 3272682

Womersley, H. & Heaslip, W. G. (1943) The Trombiculinae (Acarina) or itch-mites of the Austro-Malayan and Oriental regions. Transactions of the Royal Society of South Australia, 67, 68 - 142.

Womersley, H. (1944) Notes on and additions to the Trombiculinae and Leeuwenhoekiinae (Acarina) of Australia and New Guinea. Transactions of the Royal Society of South Australia, 68, 82 - 112.

Womersley, H. & Kohls, G. M. (1947) New genera and species of Trombiculidae from the Pacific islands. Transactions of the Royal Society of South Australia, 71, 3 - 12.

Womersley, H. (1952) The scrub-typhus and scrub-itch mites (Trombiculidae, Acarina) of the Asiatic-Pacific region. Records of the South Australian Museum, 10, 1 - 435.

Womersley, H. & Audy, J. R. (1957) Malaysian Parasites-XXVII. The Trombiculidae (Acarina) of the Asiatic-Pacific Region: A revised and annotated list of the species in Womersley, 1952, with descriptions of larvae and nymphs. Studies from the Institute for Medical Research, Federation of Malaya, 28, 231 - 296.

Yu, Z. Z., Yang, G. R. & Chen, R. H. (1980) Two new species of the genus Ascoschoengastia from Yunnan, China (Acarina: Trombiculidae). Acta Zootaxonomica Sinica, 5, 150 - 153. [in Chinese]

Yu, Z., Yang, G., Zhang, T., Zi, D. & Hu, G. (1981) Descriptions of five new species of Leptotrombidium from China (Acarina: Trombiculidae). Acta Zootaxonomica Sinica, 6, 149 - 156. [in Chinese]

Yu, Z. Z., Chen, R. H. & Lin, J. B. (1996) Five new species of chigger mites from Yunnan Province, China (Acari: Trombiculidae). Acta Zootaxonomica Sinica, 21, 430 - 437. [in both Chinese and English]

Zhao, S., Tang, F. & Mo, G. (1986) A study of the chigger mites and their hosts from Xisha Archipelago, China (Acarina: Trombiculidae). Acta Entomologica Sinica, 29, 96 - 100. [in Chinese]