Lindsaea quadrangularis Raddi subsp. terminalis K.U.Kramer

Matos, Fernando B., Bohn, Amabily & Labiak, Paulo H., 2020, The ferns and lycophytes of Reserva Natural Guaricica, Antonina, Paraná, Brazil, Check List 16 (1), pp. 183-206 : 194-196

Lindsaea quadrangularis Raddi subsp. terminalis K.U.Kramer

Figure 5A

Examined material. BRAZIL • 2 specimens; Trilha do Gervásio ; 25°14′39″S, 048°39′59″W, alt. 340 m; 27 May 2006; F.B. Matos & M.C. Gomes 1179 leg.; MBM330188, UPCB6065 • 1 specimen; Trilha Esborracha Faisqueira ; 25°23′37″S, 048°40′52″W; alt. 15 m; 12 Jun. 2019; F.B. Matos et al. 2648 leg.; UPCB44506 .

Identification. Characterized by creeping, scaly stems, the petioles reddish-brown to blackish or atropurpureous, subterete at the extreme base, otherwise quadrangular, the blades 2-pinnate, and the pinnules dimidiate. Sori submarginal, continuous along the upper margins, indusiate (indusia open outwardly). Differs from most Lindsaea species by the dark (vs stramineous) axes. Lindsaea divaricata Klotzsch also has dark axes, but differs by the presence of two lateral spreading wings on the adaxial side of rachises.

Distribution and habitat. Paraguay and Brazil. Terrestrial in forests.

Order Polypodiales (suborder Pteridineae J.Prado & Schuettp. ), Pteridaceae E.D.M.Kirchn.