Friesodielsia species

Leeratiwong, Charan, Karapan, Sunate, Satthaphorn, Jiratthi & Johnson, David M., 2023, Two new species of Friesodielsia (Annonaceae) from Peninsular Thailand, Phytotaxa 589 (1), pp. 73-82 : 80-81

Key to the Friesodielsia species of Thailand ( (modified from Leeratiwong et al., 2022)

1. Outer petals appressed until anthesis, then separating slightly, triquetrous apically..........................................................................2

- Outer petals separating widely in early bud stage, flat apically .........................................................................................................9

2. Twigs persistently hispid, pedicels 59–95 mm long, sepals with 1 vein, leaf base irregularly cordate ................................. F. filipes

- Twigs tomentose, pubescent or glabrous, pedicels less than 59 mm long, sepals obscurely to strongely 5–7 veined or without visible veins, leaf base regularly cordate, rounded or broadly cuneate..............................................................................................3

3. Sepals 7.0–17.0 mm long ...................................................................................................................................................................4

- Sepals 2.5–6.0 mm long .....................................................................................................................................................................6

4. Pedicellar bract apical, leaf apex rounded or truncate-retuse, rarely with a short mucro ca. 2 mm long., outer petals lanceolate, inner half the length of the outer petals ....................................................................................................................................... F. calycina

- Pedicellar bract at pedicel midpoint or basal, leaf apex acute, acuminate, ormucronate, outer petals linear-oblong or linearlanceolate, inner petal greater than half the length of the outer petals ...............................................................................................5

5. Stem densely pubescent, leaves glaucous abaxially, floral pedicels 6–10 mm long, with cordate to reniform bracts, sepals with 5 distinct veins, outer petals with a raised abaxial midrib, carpels 7–15 .................................................................................. F. affinis

- Stem sparsely pubescent, leaves shiny abaxially, floral pedicels 10–15 mm long with broadly ovate bracts, sepals with 5–7 obscure veins, outer petals without raised abaxial midrib, carpels ca. 30 .......................................................................... F. chalermgliniana

6. Pedicels of flower/fruit ≥ 14 mm long ................................................................................................................................................7

- Pedicels of flower/fruit <14 mm long ................................................................................................................................................8

7. Twigs glabrous or sparsely hairy, leaves 5–12 cm wide, outer petals linear, inner petal length greater than half the length of the outer petals............................................................................................................................................................................. F. glauca

- Twigs densely hairy, leaves less than 5 cm wide, outer petals lanceolate to ovate-lanceolate, inner petal length greater than half the length of the outer petals ............................................................................................................................................... F. betongensis

8. Outer petals linear-lanceolate, acute, sepal apex acute/acuminate, leaf blades 3.5–7.5 mm wide, apex acuminate/caudate, acumen (3–) 7–20 mm long ................................................................................................................................................... F. khaoluangensis

- Outer petals lanceolate, obtuse, sepal apex rounded, leaf blades 7.5–15.0 cm wide, apex obtuse to rounded, occasionally emarginate or mucronate........................................................................................................................................................................ F. latifolia

9. Twig hairs persistently appressed, often glabrate proximal to first fully expanded leaves on specimen.........................................10

- Twig hairs at first appressed, soon becoming erect to spreading, twigs always pubescent proximal to first fully expanded leaves on a specimen ........................................................................................................................................................................................12

10. Bracts 1.5–3.5 mm long, shorter than pedicel, monocarp stipes 3–5 mm long................................................................... F. discolor

- Bracts 4.0–9.0 mm long, longer than pedicel, monocarp stipes 5–9 mm long ................................................................................11

11. Leaves glaucous abaxially, inner petals lobed at midpoint on both sides ....................................................................... F. unonifolia

- Leaves shiny abaxially, inner petals without lobes at midpoint ............................................................................................ F. glabra

12. Leaves spreading-pubescent below ..................................................................................................................................................13

- Leaves appressed-pubescent to glabrate below................................................................................................................................14

13. Monocarps ellipsoid to oblongoid, (8.0–)10.0–14.0 mm wide, sepals (5.0–)6.5–10.0 mm long, outer petals 40–70 Í 13–35 mm .... ................................................................................................................................................................................................. F. kingii

- Monocarps ovoid, 6.5–7.0 mm wide, sepals 4–5 mm long, outer petals (19–)27–31 Í 7–9 mm .................................... F. desmoides

14. Sepals ≤ 10 mm long .........................................................................................................................................................................15

- Sepals> 10 mm long .........................................................................................................................................................................16

15. Inner petals 8–13 mm long, ovate-lanceolate to ovate, with marginal lobes, monocarp stipes 3–5 mm long............ F. brevistipitata

- Inner petals (11–) 15–25 mm long, narrowly lanceolate to lanceolate, lacking marginal lobes, monocarp stipes 5–10 mm long....... ........................................................................................................................................................................................ F. longipetala

16. Sepals gradually tapering to a blunt-acuminate apex, outer petals 6–12 mm wide, inner petals 8–10 mm long, petioles grooved adaxially .................................................................................................................................................................... F. songkhlaensis

- Sepals rounded to obtuse or acute apex, outer petals 11–32 mm wide, inner petals 10–25 mm long, petioles flat adaxially.........17

17. Monocarp stipes 2–5 mm long, swollen apically ....................................................................................................... F. phanganensis

- Monocarp stipes 5–10 mm long, uniformly thick ............................................................................................................................18

18. Outer petals (50–) 80–140 mm long, carpels 10–14, monocarps sparsely pubescent..................................................... F. longipetala

- Outer petals 50–80 mm long, carpels 14–24, monocarps densely pubescent ............................................................. F. macrosepala