Omma stanleyi Newman, 1839

Escalona, Hermes E., Lawrence, John F. & Ślipiński, Adam, 2020, The extant species of the genus Omma Newman and description of Beutelius gen. nov. (Coleoptera: Archostemata: Ommatidae: Ommatinae), Zootaxa 4728 (4), pp. 547-574 : 551-552

Omma stanleyi Newman, 1839

( Figs 1–3 , 14–18, 21 , 39–43 , 51–55 , 58–60, 63 , 64–66 , 69–70 , 74 , 76–78, 80–81 , 90 )

Omma stanleyi Newman 1839: 304 . Neboiss (1968, 1989); Lawrence (1999).

Diagnosis. Body clothed with moderately short and decumbent yellowish setae. Head elongate, 1.3 times as long as wide. Frontoclypeus slightly concave. Prothorax slightly transverse, 0.7 times as long as wide. Elytra 2.0 times as long as wide (EL/EW), surface without raised carinae or tubercles ( Figs 2 , 59 ); epipleura narrow ( Fig. 58 ). Aedeagus with parameres longer than penis ( Fig. 90 ).

Description. Length: ♂ 14–20 mm, ♀ 14.4–27.5 mm. Body 1.3 times as long as wide (BL/EW), integument dark brownish, clothed with moderately short and decumbent yellowish setae. Head elongate, 1.3 times as long as wide, surface with uniform setae but longer dorsally around the eyes and temple. Frontoclypeus slightly concave. Labrum transverse with more or less truncate apex bordered with pointed tubercles. Gulamentum with setae converging to the disc. Mentum as long as wide, slightly narrowing towards the apex, surface concave. Antennae not extending beyond base of prothorax; antennomere (♂) length/width ratios, 1: 1.0, 2: 0.8, 3: 2.5, 4: 1.4, 5: 1.2, 6: 1.3, 7: 1.3, 8: 1.2, 9: 1.2, 10: 1.3, 11: 1.7; antennomere 5 with one sensory area, antennomeres 6–9 with two sensory areas and antennomere 11 with one large sensory area. Apical labial and maxillary palpomeres expanded and securiform.

Prothorax with uniform setae, slightly transverse, 0.7 times as long as wide, pronotal disc slightly convex with a transverse depression between posterior impressions; notopleural suture complete, sternopleural suture absent; prosternal process incomplete and short. Elytra surface without carinae, widest around mid-length. Outer surface of protibia in male often with small setose excavation at basal third to fourth (fig. 9 in Lawrence 1999, see comments). Protarsus (♂) 0.7 times as long as protibia (TAL/TIL). Mesotarsus 1.2 times as long mesotibia (TAL/TIL). Meta- tarsus 1.9 times as long as metatibia (TAL/TIL). Abdominal ventrite 5 posterior edge partially margined. Aedeagus with penis narrowing to apex, parameres with apical oval notch and translucent apical portion extended forward well beyond apex of penis ( Fig. 90 ).

Comments. The size and presence of the male protibial dimorphism is variable, it can be absent (e.g., Biggenden, nr. Rifle Range) or present in one leg only (e.g., Macedon, VIC), and in some the setose excavation is reduced to only 1–2 setae. The gut contents of one specimen (Biggenden, nr. Rifle Range) included abundant items resembling pollen or spores.

Material examined. Holotype: Stanleyi Melly, Nov. Holl., NG Omma Newman , Typus ( NHMG). NEW SOUTH WALES: Bogan River , 1.i.1931 (1 CAS). Brewarrina , 1916, W.W. Froggatt (1 ♀ ANIC). Cootamundra , 20135 (1 ♀ SAMA-25-038600). Mogriguy (2 AM). Mudgee (1 CAS). Oatley , xii.1943 (2 CAS). Ponto Falls via Wellington , 12.xii.1977, F.J. Grant-Taylor & D. R. Smith (1 ♀ ANIC). New records: Muswellbrook, E.W. Ferguson Coll. (1 ♂ ANIC). Rope’s Creek , ex University of Sydney , MacLeay Museum (1 ♀ ANIC); same locality (1 ♀ SAMA-25-038597); same locality, F.E. Wilson Coll. (1 ♀ VIC). Penrith (2 ♂♂, 1 ♀ AM). Mogriguy , 1–5.i.1979, R. H. Mulder (2 ♀♀ AM). 5.4 KM along “MIDGERY” Station road from junction with Gwydir Hwy., 29.38.52 148.20.21, Eucalyptus populnea patch, 21.xi–11.xii.1999, F. Christie, P. Flemons, M. Elliot, pit trap (1 ♀ AM). Nov Holl , Fry Coll. 1905-100 (1 ♀ MNHL). No locality (1 ♀ VIC). QUEENSLAND: Biggenden , nr. Rifle Range , 5.i.1974, H. Frauca (1 ♂, ANIC). Braemar State Forest , via Kogan (27.11 S, 148.03 E), 15–19.x.1979, G.B. Monte- ith (1, QMB). Bungle Creek , 26 km N of Roma, 20.i.1984, A. Hiller (1, DPIM). Duaringa (1 ♂ SAMA-25-038591). Edungalba, i.1964, J.G. Brooks (1 ♂ ANIC); same locality, 24.i.1974, E. Adams (1, DPIM). Milmerran, 3.iii.1964, J.G. Brooks (1, ANIC). Wingham, 1898 (1, ANIC). New Records: Dalby, M.F. R. Hobler (1 ♀ AM, 1 ♂ SAMA- 25-038595, 1 ♀ SAMA-25-038594). Gayndah (1 ♂ AM). Warwick 1952, E.B. Miller (1 ♀ AM); same locality, oct.1940, E. Smith (1 ♂, 1 ♀ VIC). Clermont, Dr. K.K. Spence (2 ♀♀ ANIC); same locality, x.29, Dr. K.K. Spence (6 ♀♀ AM); same locality, 22.x.1929, K. Spence (1 ♂ ANIC dissected); same locality, Dr. K.K. Spence, F.E. Wilson Collection (1 ♂ VIC). Mt. Tamborine , 7.i.1963, B.P. More (1 ♂ ANIC). Ex. Coll. Nat. Mus., 3112 (1 ♀ ANIC). G. Lewis Coll. BM 1910-248 (1 ♂ MNHL). C. French’s Coll., 5.xi.1908 (1 ♀ VIC); same data, 3.xi.1908 (1 ♀ VIC). No data (1 ♂ AM, 1 ♀ SAMA-25-038588, 1 ♂ dissected, SAMA-25-038590). SOUTH AUSTRALIA: Dawson,

x.1878, D. Sharp Coll. B.M. 1932-116 (1 ♀ ANIC) ; same locality, D. Sharp Coll. BM 1932–116 (1 ♀ MNHL) . New records: Nuriootpa (1 ♂ SAMA-25-038586). VICTORIA: Lillimur , 12.iii.1888, L. Molinaux (1 ♀ SAMA-25- 038592). Melbourne, Niechoff (1 ZISP) . New records: Inglewood (1 ♀ VIC, Figs 2–3 ). Melbourne, 4826 (1 ♂, 1 ♀ MNHL) . Ballarat, C. Oke (1 ♀ VIC) . Macedon, C. Oke (1 ♂ VIC). UNKOWN STATE: New records: Griffith Collection, id. A.M. Lea (1 ♂ SAMA-25-038598, 1 ♀ SAMA-25-038599). Sharp Coll. 1905-313 (1 ♀ MNHL) . Pascoe Coll. 93-60 (1 ♂ MNHL) . 21505, Nov. Holl., Bor, Fry Coll. 1905-100 (1 ♀ MNHL) . F.E. Wilson Coll. (1 ♂ VIC) . Chilerum? (1 ♀ VIC) . No locality (1 ♂ SAMA-25-038596, 1 ♂ SAMA-25-038593, 1 ♂ SAMA-25-038587, 1 ♀ SAMA-25-038589, 1 ♀ VIC) .

Figures

FIGURES 1–3. Omma stanleyi: 1, Natural habitat, Queensland, Australia ©Dash Huang (17 Jan 2017); 2–3, habitus: 2, dorsal (♀ 20 mm, VIC); 3, same, ventral.

FIGURES 14–21. Omma stanleyi: 14–19. Head (♀): 14, dorsal, 15, ventral, 16, lateral, 17, frontal, 18, mandible, dorsal; 19, Beutelius rutherfordi (paratype ♂): labial palp; 20, B. mastersi (♀): maxilla; 21, O. stanleyi: head, latero-ventral.

FIGURES 39–44. Omma stanleyi (♀): 39–43. 39, pronotum, 40, prothorax, ventral, 41–42, prothorax, lateral, 43, scutellum; 44, Beutelius rutherfordi, prothorax, lateral (paratype ♂).

FIGURES 51–55. Omma stanleyi (♀): 51, pterothorax, ventral; 52, pterothorax, latero-ventral; 53, pterothorax and hind legs; 54, bifid mesoventral process; 55, abdominal ventrites 1 and 2 (in part).

FIGURES 56–63. Beutelius sagitta (♂): 56, epipleuron, ventral view; B. rutherfordi (♂): 57, epipleuron, ventral view; O. stanleyi (♀) elytron: 58–60, 63. 58, ventral view with epipleuron, 59, dorsal, 60, internal, 63 dorsal; B. mastersi (♀) elytron: 61–62. 61, dorsal, 62, ventral view with epipleuron.

FIGURES 64–66. Omma stanleyi (♀) wing: 64, complete view; 65, wing base (in folded wing); 66, wing, apical field, with apical wing folding. Veinal nomenclature abbreviations: BR: radial basivenale. BM: redial basivenale. BCu: cubital basivenale. AA: anal anterior. AP: anal posterior. Cu: cubitus. C: costa. ScA: subcosta anterior. ScP: subcosta posterior. RA: radius anterior. RP: radius posterior. MP: media posterior. WC: wedge cell. BC: brachial cell. Cross-veins: rp-mp, r, cv1: mp3+4-cua, cv2: mp1+2-mp3+4, cv3: mp1+2-mp3.

FIGURES 67–70. Beutelius mastersi (♂): 67, wing (veinal nomenclature as in Fig. 64); B. sagitta (♂): 68, wing; Omma stanleyi: 69–70. 69, elytron base, 70, elytron, close detail of sutural flange.

FIGURES 71–75. Omma stanleyi and Beutelius spp. abdominal ventrites and tergites: 71, B. rutherfordi (♂, holotype); 72, B. mastersi (♀) ventrites; 73, B. sagitta (♂) ventrites; 74, O. stanleyi (♀) ventrites; 75, B. sagitta (♂), tergites.

FIGURES 76–84. Omma stanleyi (♀): 76–78. 76, foreleg, 77, midleg, 78, hind leg. Forelegs (♂): 79–84. 79, Beutelius rutherfordi; 80–81, O. stanleyi, 80, anterior view, 81, lateral view; 82–83, B. sagitta, 82, anterior view, 83, posterior view; 84, B. mastersi, anterior view.

FIGURES 85–95. Beutelius spp., mentum: 85–86. 85, B. mastersi, 86, B. rutherfordi (paratype ♂); 87, B. mastersi, mesonotum; 88, B. mastersi, pterothorax, ventral; 89, B. rutherfordi, pterothorax, ventral; aedeagus: 90–95. 90, Omma stanleyi; 91–92, B. mastersi; 93, B. reidi sp. nov.; 94, B. rutherfordi; 95, B. sagitta.