Telephanini LeConte 1861

Thomas, Michael C. & Nearns, Eugenio H., 2008, A new genus of telephanine Silvanidae (Coleoptera: Cucujoidea), with a diagnosis of the tribe and key to genera, Insecta Mundi 2008 (48), pp. 1-14 : 2-6

Tribe Telephanini LeConte 1861

Diagnosis. Within the Brontinae , all members share inverted male genitalia, a dorsal mandibular mycangium, longitudinal frontal grooves, tuberculate to spinose lateral pronotal margins, and elongate to very elongate antennal scapes. Members of Telephanini differ from members of Brontini in their posteriorly closed anterior coxal cavities, narrowly separated anterior and middle coxae, and lobed tarsomere III.

Form. Elongate, fusiform to ovate, mostly elongate, parallel-sided; size small to medium, 3-7mm in length.

Taxon/Character 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 _____________________________________________________________________________________________________________________ Cucujidae

Pediacus subglaber * 0 0 1 1 0 1 0 0 1 0 1 1 0 0 1 1

Silvanidae : Brontinae : Telephanini

Telephanus atricapillus Erichson 1 1 0 0 1 0 1 1 0 1 1 1 0 0 1 0 Aplatamus sp. 0 1 0 1 0 0 1 0 0 1 0 1 1??? Euplatamus sp. 0 1 0 1 0 0 1 1 0 1 0 1 0??? Psammoecus sp. * 1 1 0 0 0 0 1 1 0 0 1 1 0 0 1 0 Australophanus redtenbacheri (Reitter) 0 0 0 0 0 0 1 1 0 1 0 1 1??? Cryptamorphadesjardinsi (Guérin-Menéville) 0 1 0 0 0 0 0 2 0 1 0 1 0 0 0 0 Indophanusdakshinus Pal 1 1 0 0 0 0 1 1 0? 1 1???? Megapsammoecus christinae Karner 0 0 0 0 0 0 1 2 0 1 0 1 1??? Psammaechidius spinicollis Fairmaire 1 1 0 0 0 0 1 1 0 1 0 1 0???

Silvanidae : Brontinae : Brontini

Uleiota dubius (Fabricius) 0 1 0 1 0 0 0 0 0 0 1 1 0 0 0 0 Parahyliota fallax (Grouvelle) * 0 1 0 1 1 0 0 0 0 0 1 1 1 0 0 1 Macrohyliota spinicollis (Gory) 0 1 0 1 1 0 0 0 0 0 0 1 0??? Australohyliota chilensis (Blanchard) * 0 1 0 1 1 0 0 0 0 0 0 1 0 0 0? Dendrophagus cygnaei Mannerheim * 0 1 0 1 0 0 0 0 0 0 0 1 0 0 0?

Silvanidae : Silvaninae

Nausibius major Zimmermann 0 0 1 1 1 1 1 0 0 0 1 0 0???

Oryzaephilus surinamensis (L.) 0 0 1 1 1 1 1 0 0 0 1 0 0 1 1 1 Silvanus planatus Germar 0 0 1 1 1 1 1 0 0 0 1 0 0???

Ahasverus rectus (LeConte) * 0 0 1 1 1 1 1 1 0 0 1 0 0 1 1 1 Cathartus quadricollis (Guérin-Menéville) 0 0 1 1 1 1 1 1 0 0 1 0 0???

Monanus concinnulus (Walker) * 0 0 1 1 1 1 1 1 0 0 1 0 0 1 1 1 _____________________________________________________________________________________________________________________

*Notes: For Pediacus , larval characters from P. japonicus Reitter ( Klausnitzer 2001) ; for Psammoecus , larval characters from P. trimaculatus Motschulsky ( Sen Gupta and Pal 1996); for Parahyliota , larval characters from P. indica (Arrow) ( Sen Gupta and Pal 1996) ; for Australohyliota chilensis , larval characters from Cekalovic and Quezada (1972); for Dendrophagus , larval characters from D. longicornis Reitter ( Klausnitzer 2001) ; for Ahasverus , larval characters from A. advena (Waltl) ( Sen Gupta and Pal 1996) ; for Monanus concinnulus , larval characters from Sen Gupta and Pal 1996.

Surface sculpture and pubescence. Moderately to densely punctate, with pubescence usually long and conspicuous. None known with surface encrustation.

Head. Transverse, somewhat triangular in shape; frontoclypeal suture present or not; apex of clypeus subtruncate, epistome usually deeply emarginate over antennal insertions; frontal region bounded laterally by longitudinal grooves in most genera; temples slightly to markedly produced; head usually transversely impressed behind temples and abruptly constricted to form neck; gular region simple or semicircularly impressed; gular sutures present, widely separated, divergent posteriorly. Mouthparts. Mandibles stout, laterally expanded or carinate, with two apical teeth and a dorso-basally located, small to large mycangium bounded anterolaterally with a tubercle ( Fig. 11 ); prostheca and mola present. Labrum short, transverse, apparently immovable. Mentum transverse. Maxillary palps with apical palpomere simple to strongly securiform; galea broadly rounded and densely setose; lacinia with two apical teeth. Labial palps with terminal palpomere simple to securiform. Antennae. Filiform, elongate; terminal antennomeres not forming a distinct club; scape elongate to very elongate; pedicel usually shorter than antennomere III. Eyes. Moderate to large, flattened to hemispherical.

Thorax. Pronotum. Quadrate to transverse, lateral margins tuberculate to spined. Prosternum. Anterior coxal cavities narrowly separated ( Fig. 10 ) (moderately separated in Aplatamus ), closed posteriorly; intercoxal process narrow; protrochantin not exposed. Scutellum. Moderate, rounded or angulate posteriorly, usually with a distinct transverse carina or groove paralleling posterior margin ( Fig. 15 , 17 , 23, 24 ). Mesosternum. Meets metasternum in a straight to curved line, suture simple; mesocoxal cavities approximate to narrowly separated ( Fig. 10 ) (moderately separated in Aplatamus ), open laterally; closed by mesepimeron only, mesepisternum does not contribute to closure; mesepisternum apparently fused to mesepimeron and mesosternum, sutures obliterated ( Fig. 9 ); mesotrochantin not exposed. Metasternum. Transverse; discrimen present for half or more of length.

Elytra and hind wings. Elytra striatopunctate, disk usually with six or seven rows of punctures plus a scutellary striole ( Fig. 17 , 23, 24 ) (absent in Telephanus , Psammoecus , and Indophanus ); intervals flat to convex; lateral margin explanate to a greater or lesser degree, sometimes denticulate; epipleura usually complete to about apical fifth, narrow to very broad. Hind wing venation well developed ( Fig. 18- 20 ).

Legs. Usually elongate; tibial spurs present, short, subequal; tarsal formula 5-5- 5 in both sexes; tarsomere I short, oblique; II usually simple, sometimes lobed; III strongly lobed (except in Aplatamus ) or bilobed; IV very short and deeply imbedded in III; V elongate, tarsal claws simple ( Fig. 14 ).

Abdomen. Five subequal sternites, punctate, no sexual dimorphism; intercoxal process narrow; femoral lines closed.

Genitalia. Male genitalia of inverted cucujoid type. Parameres located on the ventral aspect of the median lobe, usually elongate and articulated ( Fig. 22 ), reduced in Aplatamus ( Fig. 27 ) and Megapsammoecus , absent in Australophanus ( Fig. 21 ); internal sac usually with a flagellum.

Figures

Figure 7-11. Australophanus redtenbacheri.7) Head, dorsal. 8) Prosternum. 9) Mesocoxal cavity and associated sclerites. 10) Mesosternal process. 11) mandibles, dorsal

Figure 12-15. Australophanus redtenbacheri. 12) Maxillary palp. 13) labial palp. 14) tarsus. 15) scutellum.

Figure 16-17. Australophanus redtenbacheri. 16) Head, dorsal. 17) Scutellum and base of elytra.

Figure 23-24. Scutellum and base of elytra. 23) Euplatamus sp., Bolivia. 24) Aplatamus sp., Mexico.

Figure 18-20. Hind wings. 18) Australophanus redtenbacheri. 19) Euplatamus sp., Bolivia. 20) Aplatamus sp., Mexico.

Figure 21-22. Male genitalia, A, tegmen; B, aedeagus. 21) Australophanus redtenbacheri. 22) Euplatamus sp., Bolivia.

Figure 25-27. 25) Euplatamus sp., Bolivia, head. 26-27) Aplatamus sp., Mexico. 26) Head. 27) Tegmen.