Cyclocephala carbonaria Arrow, 1911

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 147-148

Cyclocephala carbonaria Arrow, 1911

Cyclocephala carbonaria Arrow, 1911: 173-174 [original combination].

Mononidia carbonaria (Arrow) [new combination by Casey 1915: 114-115].

Cyclocephala carbonaria Arrow [revised combination by Arrow 1937b: 8, 9].

Mononidia carbonaria (Arrow) [revised combination by Bolívar y Pieltain et al. 1963: 185].

Cyclocephala carbonaria Arrow [revised combination by Endrődi 1966: 164].

syn. Mononidia carbonaria punctulata Prell, 1934: 162 [original combination].

Cyclocephala carbonaria var. punctulata (Prell) [new combination and new infrasubspecific status by Arrow 1937b: 9].

Cyclocephala carbonaria ab. punctulata (Prell) [revised infrasubspecific status by Endrődi 1964: 466].

Cyclocephala carbonaria Arrow [synonymy by Endrődi 1966: 164].

syn. Mononidia carbonaria trachypyga Prell, 1934: 162 [original combination].

Cyclocephala carbonaria Arrow [synonymy by Arrow 1937b: 9].

syn. Cyclocephala howdeni Endrődi, 1967b: 83-84 [original combination].

Cyclocephala carbonaria Arrow [synonymy by Ratcliffe 2003: 99].

Types.

Type of C. carbonaria at BMNH ( Endrődi 1966). Holotype ♂ of C. howdeni at ZMHB ( Endrődi 1967b). Location of the Prell types was not reported by Endrődi (1966).

Distribution.

BELIZE: Cayo, Stann Creek. BOLIVIA. COLOMBIA: Boyacá, Chocó, Córdoba, Cundinamarca, Santander, Valle del Cauca. COSTA RICA: Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas. ECUADOR: Guayas, Santo Domingo de los Tsáchilas. GUATEMALA: Alta Verapaz, Huehuetenango, Izabal, Petén, Zacapa. HONDURAS: Atlántida, Colón, Cortés, Olancho, Santa Bárbara, Yoro. MEXICO: Chiapas, Puebla. NICARAGUA: Chontales, Jinotega, RAA Norte, Río San Juan. PANAMA: Bocas del Toro, Chiriquí, Coclé, Colón, Former Canal Zone, Panamá, San Blas. VENEZUELA: Amazonas, Bolívar.

References.

Casey 1915, Prell 1934, Arrow 1911, 1937b, Blackwelder 1944, Bolívar y Pieltain et al. 1963, Pike et al. 1976, Endrődi 1964, 1966, 1967b, 1985a, Maes 1987, 1994, Joly 1995a, Dechambre 1991a, 1997, Ratcliffe and Morón 1997, Beath 1998, Ratcliffe 2002a, 2003, Restrepo et al. 2003, Neita-Moreno et al. 2006, Ratcliffe and Cave 2006, Pacheco-F. et al. 2008, Neita-Moreno 2011, Krajcik 2005, 2012, Breeschoten et al. 2013, Ratcliffe and Cave 2013, García-López et al. 2013, Ratcliffe et al. 2013, Yepes-Rodriguez et al. 2013, López-García et al. 2015, Gasca-Álvarez and Deloya 2016.