Lophocaterini Crowson, 1964, 1992

Kolibac, Jiri, 2013, Trogossitidae: A review of the beetle family, with a catalogue and keys, ZooKeys 366, pp. 1-194 : 121-122

Tribe Lophocaterini Crowson, 1964

Lophocaterinae Crowson, R. A. 1964a: 297 ().

Type genus:

Lophocateres Olliff, 1883

Barron, J. R. 1971: 11, 12 (syn. Lophocateridae = Peltinae ). Barron, J. R. 1975: 1119. Burakowski, B. et al. 1986: 119 ( Lophocateridae ). Kolibáč, J. 2006: 128 (diagnosis, stat. n.). Kolibáč, J. 2007a: 365. Kolibáč, J. 2010: 35. Lafer, G. Sh. 1992: 83 (key). Lawrence, J. F. & Newton, A. F., Jr. 1995: 868 ( Lophocateridae ). Lucht, W. 1998: 207 (key). Ślipiński, S. A. 1992: 442 ( Lophocaterinae )

Lycoptini Casey, 1890 (Type genus: Lycoptis Casey, 1890)

Kolibáč, J. 2006: 128 (synonymized)

Remarks.

The main issue to be addressed for Lophocaterini is their possible paraphyly in relation to Ancyronini . The whole clade (lophocaterins + ancyronins) is monophyletic but the lophocaterins might be paraphyletic (i.e., non-holophyletic in the traditional Hennigian meaning) because ancyronins can only be advanced members of more primitive lophocaterins. See also “Remarks” in the Ancyronini section. Further, more detailed study is required to resolve the question. The generic composition of Decamerini and its position within Lophocaterinae should be examined along - not, however, before an associated larva of the decamerins is known.

A key to genera

(after Kolibáč 2010)

1	Elytra with irregular punctation; lateral margins of pronotum broadly explanate, lateral edge sparsely denticulate	Eronyxa
-	Elytra regularly punctate; lateral margins of pronotum narrowly explanate, lateral edge almost entirely evenly rounded or densely denticulate	2
2	Antenna 7- or 9-segmented	3
-	Antenna 11-segmented	5
3	Antenna 7-segmented, club 1-segmented; mandible with mola	Lycoptis
-	Antenna 9-segmented, club 2- or 3-segmented; mandible without mola	4
4	Antennal club 2-segmented; mandible with prostheca near base of mandible formed by tuft of long setae; submental area lacking concave or depressed area; wing with oblong radial cell	Grynocharina
-	Antennal club 3-segmented; mandible without penicillus or prostheca; submental area concave; wing with small triangular radial cell displaced downwards	Peltonyxa
5	Lateral edge of pronotum densely denticulate; lacinia with one pigmented spine	Indopeltis
-	Lateral edge of pronotum evenly rounded or at most finely undulating; lacinia with two or three pigmented spines	6
6	Elytra with inconspicuous carinae; mola absent; ligula deeply emarginate; probably predatory. Larva: sensory appendix very short	Promanus
-	Elytra with conspicuous carinae; mola or remnant of mola present; ligula deeply or weakly emarginate. Mode of life: predatory ( Trichocateres ), herbivorous ( Lophocateres ), fungivorous ( Grynocharis ). Larva ( Lophocateres , Grynocharis ): length of sensory appendix about half or more than half that of antennal segment 3	7
7	Elytra with six carinae; tegmen without projecting phallobasic apodeme; lacinia with three pigmented, hooked spines; small species (less than 3 mm)	Lophocateres
-	Elytra with five or four distinct carinae; tegmen with projecting phallobasic apodeme; different pattern of pigmented lacinial spines; larger species (above 5 mm)	8
8	Elytra with four distinct (higher) carinae and another three to four lower carinae among them; pronotum and elytra without tufts of long hairs, with short decumbent or semi-erect pubescence only, or without conspicuous pubescence; lacinia with two pigmented, hooked spines; tibial apical spur pattern 2-2-2; larger species (about 4.5-10.5 mm)	Grynocharis
-	Elytra with only five distinct carinae; pronotum and elytra with tufts of long, yellow-orange hairs; lacinia with three pigmented spines at apex in pattern 1+2, apical spine large and hooked, two other spines much smaller; tibial apical spur pattern 1-1-1; smaller species (about 5 mm)	Trichocateres