Penniverpa festina (Coquillett)

Webb, Donald W. & Metz, Mark A., 2008, A revision of the New World genus Penniverpa Irwin and Lyneborg (Diptera: Therevidae: Therevinae), Zootaxa 1720, pp. 1-45 : 21-29

Penniverpa festina (Coquillett)

( Figs. 36–43 , 83 , 88 )

festina Coquillett (1893: 225 male /fem. desc.), Cole (1923: 35 male key, 67 notes, dist., Figs. 82 ant., 106 male genit.; 1965:351 cat.) Kröber (1913:30 listing; 1914: 38 male key) ( Psilocephala ); Metz and Irwin (2000:986 phylog.) ( Penniverpa ). Type locality United States, Florida. Lectotype male in MCZ.

Diagnosis. The male of Penniverpa festina is similar to P. i n s u l a r and is separated from P. i n s u l a r by having the tarsi dark brown, the PVL being present in the form of a narrow, basal spinous projection on the ventral lobe ( Fig. 39 ), the LGP of the gonostylus ( Fig. 39 ) with numerous, distinctive, thick, black macrosetae apically and the VMP positioned mediolaterally and shorter than the length of the LGP. The female of P. f e s t i n a is separated from P. i n s u l a r in having white pubescence ( Fig. 83 ) across the posterior margin of tergites 1–3 and posterolaterally on tergites 5–6; furca ( Fig. 43 ) oblong, sides constricted.

Redescription of MALE. Variation (n=10).

Body length 5.1–6.8, 6.1 mm.

Head. Length 0.70–0.90, 0.78 mm. Ommatidia of equal size. Frons pubescence white, dense (occasionally dorsal half black in narrow dorsolateral triangle separated medially); setae absent. Antenna dark brown, pubescence on scape and pedicel gray (occasionally pedicel and first flagellomere dark yellow); antenna/head length 0.67–0.81, 0.74; scape length 0.13–0.16, 0.14 mm, width 0.08–0.10, 0.09 mm, length/width 1.3–2.0, 1.6, scape/pedicel length 1.5–2.0, 1.8, scape/flagellum width 0.7–0.9, 0.9, setae black, short, sparse; pedicel length 0.07–0.10, 0.08 mm, length/width 0.7–1.0, 0.9; flagellum length 0.30–0.40, 0.36 mm, width 0.10–0.11, 0.11 mm, length/width 3.0–3.6, 3.4, flagellum/scape length 2.3–2.9, 2.5. Maxillary palpus dark brown; length 0.28–0.38, 0.32 mm, length/width 3.2–6.3, 5.2; setae dark brown. Genal setae white. Occipital setae white, absent dorsomedially; macrosetae in single row extending ventrally from lateral end of postocular macrosetae.

Thorax. Macrosetae 2–3 np, 1–2 sa. Mesonotum brownish gray; vittae dark brown, pubescence gray (occasionally vittae indistinguishable); setae black, short, erect to suberect intermixed with white, lanceolate, appressed setae. Pleuron pubescence gray; setae white on propleuron, anepisternum, katepisternum, laterotergite, and scutellum. Wing. Membrane pale smoky brown; veins brown; length 4.8–5.6, 5.2 mm, width 1.5– 2.0, 1.8 mm, length/width 2.6–3.2, 2.9, pterostigma pale brown, narrow. Halter pale brown. Legs. Coxal setae white; hind coxa with anterior papillate knob. Femora dark reddish brown, pubescence white; ventral and dorsal setae white; av 0–1:0:1, pv 0:0:2–6. Tibiae dark yellow to pale brown, apex occasionally darker. Tarsi pale brown to brown.

Abdomen. Tergite 1 dark brown, pubescence white, dense posteriorly; tergite 2 dull orange with dark brown anteromedial area, pubescence white, dense; tergites 3–8 dull orange, pubescence white, dense; dorsal setae white, elongate, appressed, denser across posterior margins. Terminalia (MEI 128494). Dark brown. Tergite 8 ( Fig. 36 ), posterior margin deeply emarginate; setae yellow with scattered dark reddish brown setae. Sternite 8 ( Fig. 37 ), setae dark reddish brown, absent medially on posterior margin. Epandrium ( Fig. 36 ), setae dark reddish brown, thicker than setae on tergite 8. Cercus ( Fig. 36 ), ending basal to apex of hypoproct. Hypoproct ( Fig. 36 ), posterior margin rounded, ending distal to apex of cercus. Gonocoxite ventral view ( Fig. 37 ), setae dark reddish brown with scattered dark reddish brown macrosetae; lateral view ( Fig. 38 ) globular with elongate inner gonocoxal process; ventral lobe ( Fig. 39 ), PVL in form of short, basal, spinous projection. Gonostylus ( Fig. 39 ), LGP oblong, apical macrosetae black, coarse, VMP elongate. Aedeagus with dorsal apodeme ( Fig. 40 ), anterior margin truncate; ventral apodeme ( Fig. 41 ) ending at anterior margin of dorsal apodeme; distiphallus dorsal view ( Fig. 40 ) constricted basally, elongate, oval, tapered posteriorly with spinous dorsal projections, ventral view ( Fig. 41 ) AAP with apical spinous projections, lateral view ( Fig. 42 ), extended posteriorly or as a broad curve ventrally.

FEMALE. Similar to male except for following. Variation (n=10).

Body length 6.5–8.0, 7.3 mm.

Head. Length 0.76–0.92, 0.83 mm. Frons pubescence white; setae dark brown, short on dorsal two-thirds of frons. Antenna/head length 0.69–0.85, 0.76; scape length 0.14–0.20, 0.17 mm, width 0.10–0.12, 0.10 mm, length/width 1.5–1.8, 1.6, scape/pedicel length 1.8–2.4, 2.0, scape/flagellum width 0.8–1.0, 0.9; pedicel length 0.07–0.09, 0.08 mm, length/width 0.7–0.9, 0.8; flagellum length 0.34–0.42, 0.37 mm, width 0.11–0.12, 0.12 mm, length/width 2.8–3.5, 3.2, flagellum/scape length 2.0–2.4, 2.2. Maxillary palpus length 0.34–0.50, 0.38 mm, length/width 4.8–8.3, 6.1.

Thorax. Macrosetae 3 np, 1–2 sa. Wing. Length 5.3–6.3, 5.8 mm, width 1.8-2.2, 1.9 mm, length/width 2.9–3.2, 3.0. Legs. Anteroventral macrosetae 0–3:0:0–3, pv 0:0:3–6.

Abdomen. Tergites 1–3 dark brown, glossy ( Fig. 83 ), posterior fourth orange brown, narrower medially, pubescence white across posterior margin of tergites 1–3, tergite 4 orange brown, glossy, tergites 5–8 brown, pubescence white posterolaterally on tergites 5–7; dorsal setae dark brown, appressed on brown areas of tergites 1–3, white, appressed on orange brown areas of tergites 1–3; short, erect, dark brown on tergites 4–8. Terminalia (MEI 127476). Furca ( Fig. 43 ), length 0.36 mm, width 0.13 mm; anterolateral projection short.

Distribution. Penniverpa festina is distributed from Florida west to Arizona ( Fig. 88 ),

Habitats and phenology. Penniverpa festina has been collected in bald cypress swamps, hardwood hammocks, coastal dunes, and gopher tortoise burrows; on a variety of plants ( Asparagus setaceus (Kunth) Jessop , Bidens [leucantha var.] pilosa (L.) Griseb., Citrus paradisi Macfad. , C. sinensis (L.) Osbeck, Eugenia uniflora L., and Virbirnum sp.); by sweeping water hyacinth; at black lights and fluorescent lights; and in fruit fly, Malaise, McPhail, Steiner, and yellow pan traps in all months of the year except February.

Specimens examined. Type specimens. Two syntype males of Psilocephala festina Coquillett in the MCZ were examined: a male labeled “St. Augustine, Fla.” printed on white paper; second label “ Holotype No” printed on red paper; third label ” Type 7568” printed on red paper; fourth label “Collection C. W. Johnson” printed on white label. A THEREVIDAE M. E. Irwin Specimen # 140538 label printed on yellow paper has been added to this specimen. A female labeled “St. Augustine, Fla.” printed on white paper; second label “Allotype No” printed on red paper; third label “Collection C. W. Johnson” printed on white label. A THER- EVIDAE M. E. Irwin Specimen # 140539 label printed on yellow paper has been added to this specimen. The male (MEI 140538) is designated as the lectotype (MCZ). This lectotype is designated in order to fix the concept of Psilocephala festina and to ensure universal and consistent interpretation of the same. The female labeled MEI 140539 is designated as a paralectotype (MCZ). Other specimens. UNITED STATES. ARI- ZONA. Coconino County. Oak Creek Canyon [34.913, -111.726], 1829 m, –.VIII.–, Snow, F.H., 1ɗ MEI 0 80346 (SEMC). Graham County. Safford [32.834, -109.707], 14.VII.1914, Holt, E.G., 1ɗ MEI 0 90384 (MEIC). Pima County. Tucson [32.222, -110.926], 12.VII.1937, Knull, D.J. Knull, J.N., 1Ψ OSU 0 0 11344 (OSU). Santa Cruz County. 8.1 km S Patagonia, Patagonia Creek [31.467, -110.756], 27.VI.1961, Santana, F.J., 4ɗ MEI 128605–7, 128611 (FSCA), black light. FLORIDA. Avalon, 25.II.1948, Barger, 1Ψ MEI 0 0 3659 (FSCA), at Citrus sinensis ; San Lando, 25.V.1926, Cole, F. R., 1ɗ MEI 162949 (EMEC); 26.V.1926, Cole, F.R., 1ɗ MEI 162950 (EMEC); Sand Lake, 28.VI.1948, Link, O.D., 1Ψ MEI 0 0 3658 (FSCA). Alachua County. Austin Carey [Cary] Memorial Forest; [29.767, -82.199], 1–15.IV.1976, Fairchild, G.B., 1Ψ MEI 162618 (FSCA), flight trap; Gainesville [29.651, -82.325], 17–23.X.1993, Reinert, J.F., 1Ψ MEI 128146–7 (FSCA), Malaise trap; 17.VI.1924, Walker, F.W., 1Ψ MEI 0 0 3714 (FSCA); –. IV.1954, Hetrick, L.A., 1Ψ MEI 162875 (FSCA); –. V.1954, 1 Ψ MEI 162876 (FSCA); – 10.XII.1955, Patton, C.N., 1Ψ MEI 162874 (FSCA); 2.IV.1961, Weems, Jr., H.V., 1ɗ MEI 0 0 3718 (FSCA); 19.VI.1977, Belmont, R.A., 1ɗ MEI 162892, 1Ψ MEI 162575 (FSCA); 20.VI.1977, 1ɗ MEI 0 28377 (FSCA); 21.VI.1977, 1Ψ MEI 162361 (FSCA); 23.VI.1977, 8ɗ MEI 162887–91, 162893, 162895–6, 2Ψ 162293, 162886 (FSCA); 29.VI.1977, 1ɗ MEI 162894 (FSCA); 16.XI–3.XII.1979, Williams, D., 1ɗ MEI 0 79729 (FSCA), Malaise trap; 28.II.1984, Thomas, M.C., 1ɗ MEI 128489 (FSCA); 22–31.VII.1993, Reinert, J.F., 1Ψ MEI 128473 (FSCA), Malaise trap; 15–21.VIII.1993, Reinert, J.F., 1Ψ MEI 127288, 128127, 128482 128490 (FSCA), Malaise trap; 22–28. VIII.1993, 3 Ψ MEI 128145, 128474, 128481 128127, Malaise trap; 5–11. IX.1993, 4 Ψ MEI 127477, 128124, 128142, 128471 128127, Malaise trap; 10–16. X.1993, 1 Ψ MEI 128505 (FSCA), Malaise trap; 24–31. X.1993, 2 Ψ MEI 127289, 128144 (FSCA), Malaise trap; 11.VI.1992, Woodruff, R.E., 1ɗ MEI 115934 (FSCA); 14.VII.1994, 1ɗ MEI 115930 (FSCA); 21.VII.1994, 1Ψ MEI 115933 (FSCA); Gainesville, Agricultural Experiment Station [29.651, -82.325], 16.III. –, 1ɗ MEI 162959 (FSCA); Austin Carey [Cary] Forest [29.651, -82.325], 19.IX– 3.X.1975, Fairchild, G.B., 1Ψ MEI 162800 (FSCA); Austin Carey [Cary] Forest, Hatchett Creek [29.651, - 82.325], 7.IV.1976, Fairchild, G.B., 1Ψ MEI 162751 (FSCA), flight trap; Gainesville, Beville Heights [29.656, -82.4], 15.I.1986, Stange, L.A., 1ɗ MEI 128534, 1Ψ 128498 (FSCA); 15–30. IV.1986, 1 Ψ MEI 127472–3 (FSCA); 1–15. V.1986, 1 Ψ MEI 128479, 128472 (FSCA); 1–15. VI.1986, 2 Ψ MEI 128484, 128497 (FSCA); 16–30. XI.1986, 1 Ψ MEI 128478 (FSCA); Gainesville, Doyle Conner Building [29.651, -82.325], 13.X.1971, Weems, Jr., H.V., Artaud, C.R., 1Ψ MEI 162857 (FSCA), Malaise trap; 29–30.IX.1973, Weems, Jr., H.V., 1Ψ MEI 162873, 157843 (FSCA), Malaise trap; –. V.1982, Stange, L.A., 1ɗ MEI 128486 (FSCA), flight trap; Gainesville, Pierceís Homestead [29.651, -82.325], 16.IV.1976, Pierce, W.H., 1ɗ MEI 162921 (FSCA), flight trap; 18.IV.1976, 1Ψ MEI 162584 (FSCA), flight trap; 14.VI.1976, 1Ψ MEI 162354 (FSCA), flight trap; Gainesville, Pine Hill Estates [29.651, -82.325], 15.IV.1974, Weems, Jr., H.V., 1Ψ MEI 162808 (FSCA), at light; jct. SR 225 and SR 340 [346] [29.514, -82.231], 24.VII.1976, Dubose, L., 1Ψ MEI 162328 (FSCA), flight trap; Payne's [Paynes] Prairie [29.578, -82.354], 19.IV.1977, Fairchild, G.B., Weems, Jr., H.V., 1ɗ MEI 162916 (FSCA); 29.IV.1977, 1ɗ MEI 162917 (FSCA), flight trap; 9.V.1977, 1ɗ MEI 162925 (FSCA), flight trap; 30.V.1977, 1ɗ MEI 162926 (FSCA), flight trap; San Felasco Hammock [29.729, - 82.442], 29.IV.1977, Fairchild, G.B., Weems, Jr., H.V., 1Ψ MEI 162804 (FSCA), flight trap; 2.V.1977, 2Ψ MEI 162267, 162333, Malaise baited w/CO2; 23.V.1977, 1Ψ MEI 162773–4 (FSCA), flight trap; 30.V.1977, 1Ψ MEI 162764 (FSCA), flight trap; University of Florida Horticultural Experiment Station [29.767, - 82.199], 3.IV.1974, Wilkenson, F.R., 1Ψ MEI 162870 (FSCA), hardwood forest; 29.IV.1977, Fairchild, G.B., Weems, Jr., H.V., 1Ψ MEI 162805 (FSCA), flight trap; 5.V.1977, 1Ψ MEI 162806 (FSCA), flight trap; 9.V.1977, 1Ψ MEI 162765 (FSCA), flight trap. Brevard County. Cocoa [28.386, -80.742], –. VII.1944, Bohart, G.E., 1ɗ MEI 112494 (USNM); –. VII.1944, Bohart, G.E., 1ɗ MEI 112498 (USNM); Georgiana [28.288, -80.675], date?, Wittfeld, W., 2Ψ MEI 112464, 112501 (USNM); Palm Bay [28.035, -80.605], 16.IV.1980, Smith, F.H., 1Ψ MEI 161163 (FSCA); Merritt Island [28.357, -80.705], 14.XI.1993, Roberts, L. M., 1Ψ UCF 0 55807 (UCFC), flourescent lights. Broward County. Fort Lauderdale [26.122, -80.144], 24.VIII.1988, Phillips, T.L., 1ɗ MEI 128483 (MEIC); Hollywood [26.011, -80.145], 18.IV.1963, Rauch, P., 1ɗ MEI 157823 (MEIC). Charlotte County. 1.6 km W Hwy 31, 3.2 km N Hw [74, 26.975, -81.78], 24.IV.1989, Maynard, L.S., 1ɗ MEI 128492, 1Ψ 128493 (FSCA), in copula. Citrus County. 20.III.1957, Weems, Jr., H.V., 1ɗ MEI 0 0 3722 (FSCA). Dade County. 18.IX.1957, Denmark, H.A., 1ɗ MEI 112455 (USNM); Hialeah [25.857, -80.278], 22.VIII.1964, Stegmaier, C.E., 1ɗ MEI 162961 (FSCA), sweepnetting grasses. De Soto County. Arcadia (Arcadia Municipal Airport) [27.192, -81.837], 1.IV.1972, Miller, Kurczewski, 27ɗ MEI 112507 –8, 112510 –18, 112522 –37, 8Ψ 112538 –45 (USNM); Arcadia [27.216, -81.859], 29.III.1972, Kurczewski, F.E., Miller, R.C., 1Ψ MEI 112546 (MEIC); 30.III.1972, 3ɗ MEI 112519 –21, 1Ψ MEI 112509 (USNM). Duval County. Hanna Park Campground [30.332, -81.656], 29.V.1977, Weems, Jr., H.V., 1Ψ MEI 162807 (FSCA), hardwood hammock; Jacksonville [30.332, -81.656], 26.VI.1957, Thompson, P.H., 1ɗ MEI 112454 (USNM). Hardee County. Lemon Grove [27.558, -81.678], 1.V.1963, Rhodes, R.H., 1Ψ MEI 0 0 3651 (FSCA), Steiner trap; Bowling Green [27.638, -81.824], 21.I.1961, Vild, R.E., 1ɗ MEI 0 0 3706 (FSCA), Steiner trap; Buchanan [27.412, -81.793], 10.VI.1968, Brown, V.G., 1ɗ MEI 162962 (FSCA). Highlands County. 12 km S Lake Placid, Archbold Biological Station [27.188, -81.338], 15.IV.1989, Maier, C.T., 1ɗ MEI 111955, 1Ψ 111952 (CAES); 16.IV.1989, 1ɗ MEI 111957, 3Ψ 111951, 111953 –4 (CAES); 17.IV.1989, 1ɗ MEI 111956 (CAES); Archbold Biological Station, 12.III.1956, Frost, S.W., 1ɗ MEI 162948 (FSCA); 29.III.1961, 1Ψ MEI 112466 (USNM); 26.I.1962, 1ɗ MEI 112465 (USNM); 14.IV.1963, Rozen, J.G. Rozen, B.L., 1ɗ MEI 162941, 1Ψ 0 76035 (AMNH); 17.III.1964, Frost, S.W., 1Ψ MEI 161170 (FSCA); 21.III.1964, 1Ψ MEI 162803 (FSCA); 26.III.1964, 1Ψ MEI 162782 (FSCA); 9.X.1964, Arnaud, Jr., P.H., 1ɗ MEI 156075 (CSCA); 12.X.1964, 1Ψ MEI 156084 (CSCA); 2.III.1967, Frost, S.W., 1ɗ MEI 162947 (FSCA); 21.III.1969, Pechuman, L.L., 1Ψ MEI 162862 (CUIC); 29.III.1969, 1Ψ MEI 162854 (CUIC); 7.IV.1969, 1Ψ MEI 162864 (CUIC); 10.IV.1969, 1Ψ MEI 162863 (CUIC); 17.IV.1969, 1Ψ MEI 162865 (CUIC); 13.II.1973, Kurczewski, F.E., Miller, R. C., 1ɗ MEI 0 75756 (CUIC); 13.II.1973, 1Ψ MEI 0 75755 (CUIC); 31.III.1976, Berenbaum, M., 1Ψ MEI 0 75561 (CUIC); 25.III.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 4Ψ MEI 162452, 162302, 162458, 162600 (FSCA), flight trap; 6.IV.1978, 6Ψ MEI 162317, 162370, 162379, 162451, 162540, 162608 (FSCA), flight trap; 7.IV.1978, 1Ψ MEI 162443 (FSCA), flight trap; 8.IV.1978, 2ɗ MEI 162878, 162880, 2Ψ 162290, 162569 (FSCA), flight trap; 9.IV.1978, 5Ψ MEI 162316, 162375, 162488, 162546, 162548 (FSCA), flight trap; 11.IV.1978, 3Ψ MEI 162405, 162473, 162487 (FSCA), flight trap; 12.IV.1978, 1Ψ MEI 162533 (FSCA), flight trap; 13.IV.1978, 1ɗ MEI 162897, 3Ψ MEI 162372, 162393, 162513 (FSCA), flight trap; 14.IV.1978, Weems, Jr., H.V., Halkin, S., 1Ψ MEI 162541 (FSCA), flight trap; 14–15.IV.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 2ɗ MEI 162885, 162902, 5Ψ MEI 162417–8, 162474, 162550, 162630 (FSCA), flight trap; 17–18. IV.1978, 5 Ψ MEI 162253, 162358, 162424, 162542, 162561 (FSCA), flight trap; 19–20. IV.1978, 1 ɗ MEI 162898, 5Ψ MEI 159745, 162221, 162416, 162627, 162632 (FSCA), flight trap; 1.V.1978, Weems, Jr., H.V., 2Ψ MEI 162499, 162568 (FSCA), flight trap; 31.IV.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 6Ψ MEI 162277, 162319, 162376, 162384, 162492, 162506 (FSCA), flight trap; 11.V.1978, Weems, Jr., H.V., Klein, L.K., 4Ψ MEI 162229, 162411, 162468, 162567 (FSCA); 16.V.1978, 2Ψ MEI 162248, 162377 (FSCA), flight trap; 24.V.1978, 1Ψ MEI 162442 (FSCA), flight trap; 25.V.1978, 1ɗ MEI 162943 (FSCA), flight trap; 26.V.1978, 2Ψ MEI 162427, 162619 (FSCA), flight trap; 28–29. V.1978, 2 Ψ MEI 162250, 162301 (FSCA), flight trap; 2.VI.1978, 1ɗ MEI 162927, 3Ψ 162761–3 (FSCA), flight trap; 3.VI.1978, 2Ψ MEI 157842, 159977 (FSCA), flight trap; 4.VI.1978, 2Ψ MEI 162296, 162607 (FSCA), flight trap; 5.VI.1978, 1ɗ MEI 162954, 3Ψ 159794, 162260, 162429 (FSCA), flight trap; 10–11. VI.1978, 3 ɗ MEI 162882–4, 4Ψ 162315, 162355, 162362, 162521 (FSCA), flight trap; 18.IX.1978, Weems, Jr., H.V., Chance, S.J., 1Ψ MEI 162544 (FSCA), flight trap; 2.X.1978, 1Ψ MEI 162380 (FSCA), flight trap; 5.X.1978, 1Ψ MEI 162324 (FSCA), flight trap; 6–8. X.1978, 1 Ψ MEI 162496 (FSCA), flight trap; 10.X.1978, 1Ψ MEI 162616 (FSCA), flight trap; 11.X.1978, 1Ψ MEI 162270 (FSCA), flight trap; 12.X.1978, 2Ψ MEI 162433, 162535 (FSCA), flight trap; 13.X.1978, 1ɗ MEI 162938 (FSCA), flight trap; 14–18.X.1978, Weems, Jr., H.V., Lohrer, F.E., 2Ψ MEI 162336, 162440 (FSCA), flight trap; 27–29.X.1978, Weems, Jr., H.V., Chance, S.J., 1Ψ MEI 162366 (FSCA), flight trap; 31.X.1978, 1Ψ MEI 162285 (FSCA), flight trap; 10.XI.1978, 2Ψ MEI 162462, 162472 (FSCA), flight trap; 23.XI.1978, 2Ψ MEI 162397, 162408 (FSCA), flight trap; 26.XI.1978, Weems, Jr., H.V., Halkin, S., 1Ψ MEI 162374 (FSCA), flight trap; 8.XII.1978, Chance, S.J., 1Ψ MEI 162771 (FSCA), flight trap; 13.XII.1978, Weems, Jr., H.V., Chance, S.J., 1ɗ MEI 162877, 1Ψ 162625 (FSCA), flight trap; 15.XII.1978, 1Ψ MEI 162556 (FSCA), flight trap; 19– 20.XII.1978, Weems, Jr., H.V., Halkin, S., 2Ψ MEI 162522, 162621 (FSCA), flight trap; 2.III.1979, 1Ψ MEI 162402 (FSCA), flight trap; 11–13. III.1979, 1 Ψ MEI 162438 (FSCA), flight trap; 16–19. III.1979, 1 Ψ MEI 162292 (FSCA), flight trap; 19.III.1979, 2Ψ MEI 162387, 162626 (FSCA), flight trap; 18.IV.1979, 1Ψ MEI 157841 (FSCA), flight trap; 15.V.1979, 1Ψ MEI 162758 (FSCA), flight trap; 17.V.1979, 1Ψ MEI 162787 (FSCA), flight trap; 18–21. V.1979, 1 Ψ MEI 162784 (FSCA), flight trap; 21–22. V.1979, 1 Ψ MEI 162786 (FSCA), flight trap; 25.VI.1979, Weems, Jr., H.V., Webber, T.A., 2Ψ MEI 162752, 162759 (FSCA), flight trap; 29.VI.1979, 1Ψ MEI 162789 (FSCA), flight trap; 7–9.VII.1979, Weems, Jr., H.V., Harris, C.W., 1Ψ MEI 162753 (FSCA), flight trap; 12.VII.1979, Weems, Jr., H.V., Webber, T.A., 1Ψ MEI 162791 (FSCA), flight trap; 14–15. VII.1979, 1 Ψ MEI 162785 (FSCA), flight trap; 20–22.VII.1979, Weems, Jr., H.V., Harris, C.W., 1Ψ MEI 162788 (FSCA), flight trap; 24.VII.1979, Weems, Jr., H.V., Webber, T.A., 1Ψ MEI 162781 (FSCA), flight trap; 10–12. VIII.1979, 1 Ψ MEI 162769 (FSCA), flight trap; 24.VIII.1979, 1ɗ MEI 162924, 15– 16. IX.1979, 1 Ψ MEI 162757 (FSCA), flight trap; 21–24. IX.1979, 1 Ψ MEI 162755 (FSCA), flight trap; 31.X.1979, 1Ψ MEI 162772 (FSCA), flight trap; 1.XI.1979, 1Ψ MEI 162776 (FSCA), flight trap; 8.XI.1979, 1Ψ MEI 162779 (FSCA), flight trap; 9.XI.1979, 2Ψ MEI 162777–8 (FSCA), flight trap; 16.XI.1979, 1Ψ MEI 162780 (FSCA), flight trap; 19.XI.1979, 1Ψ MEI 162766 (FSCA), flight trap; 1.II.1980, Weems, Jr., H.V., Carrel, J. E., 1ɗ MEI 162922 (FSCA), flight trap; 21.II.1980, 1ɗ MEI 162923 (FSCA), flight trap; 28– 31. III.1980, 1 Ψ MEI 127474 (FSCA), flight trap; 4.IV.1980, 1Ψ MEI 162768 (FSCA), flight trap; 16– 17. IV.1980, 1 ɗ MEI 162904, 1Ψ MEI 162801 (FSCA), flight trap; 18–20. V.1980, 2 Ψ MEI 162767, 162812 (FSCA), flight trap; 30.IX–2.X.1980, Lampert, Jr., L.L., Weems, Jr., H.V., 1Ψ MEI 128137 (FSCA), flight trap; 20.IV.1982, Brastala, E.M., 1Ψ MEI 0 49276 (PURC); 18.III.1987, Bergson, C.S., 1ɗ MEI 162942 (MEIC), light trap; 14–17.IV.1989, Marshall, S.A., 2Ψ MEI 073586–7 (DEBU); 15–16.IV.1989, Norrbom, A.L., 1Ψ MEI 162347 (USNM); 11.IV.1991, Davis Jr., L.R., 2Ψ MEI 127475–6 (FSCA); ca. 12.9 km SW Lake Placid, ca. 9.7 km W. jct. Hwy 27 on Hwy 70 [27.167, -81.449], 16.IV.1989, Nelson, C.R., Maier, C., Fisher, E.M., Bullington, S.W., 4ɗ MEI 0 80241, 0 80243, 080245 –6 (CAS); L. Placid, Archbold Biological Station [27.188, -81.338], 5.XI.1965, Dawson, R.W., 2Ψ MEI 157821–2 (MEIC); 5.III.1966, 1ɗ MEI 157824 (MEIC); Lake Placid [27.293, -81.363], 1.XII.1958, Frost, S.W., 1ɗ MEI 112504 (USNM); Lake Placid, 12.9 km S Archbold Biological Station, 28.VIII.1982, Deyrup, M., 1ɗ MEI 162928, 1Ψ MEI 162783 (FSCA); Lake Placid, Archbold Biological Station [27.297, -81.37], 11.XI.1965, Dawson, R.W., 1ɗ MEI 162957 (WSU); 21.III.1966, 1ɗ MEI 108726 (WSU); 30.III.1966, 1ɗ MEI 162956 (WSU); 8.IV.1966, 1ɗ MEI 108725 (WSU); 10.IV.1966, 1ɗ MEI 108729 (WSU); 11.IV.1966, 1ɗ MEI 108735 (WSU); 8.VI.1966, Silberglied, R., 1Ψ MEI 162866 (CUIC), black light trap; 9.IV.1971, Lavallee, A.G., 1Ψ MEI 162760 (UGCA); 22.III.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 2Ψ MEI 162401, 162592 (FSCA); 23.III.1978, 9Ψ MEI 162254, 162303, 162421, 162431, 162437, 162450, 162532, 162571, 162596 (FSCA), flight trap; 24.III.1978, 6Ψ MEI 162268, 162274, 162309, 162341, 162349, 162464 (FSCA), flight trap; 3–4. IV.1978, 1 Ψ MEI 162775 (FSCA), flight trap; 16.IV.1978, Weems, Jr., H.V., Halkin, S., 1Ψ MEI 162565 (FSCA), flight trap; Weems, Jr., H.V., Lampert, Jr., L.L., 2Ψ MEI 162555, 162582 (FSCA), flight trap; 2–4.II.1979, Weems, Jr., H.V., Halkin, S., 1Ψ MEI 162310 (FSCA), flight trap; 28.II.1979, 1ɗ MEI 162881 (FSCA), flight trap; 2– 4. III.1979, 1 Ψ MEI 162545 (FSCA), flight trap; 5.III.1979, 1Ψ MEI 162615 (FSCA), flight trap; 9.III.1979, 1Ψ MEI 162449 (FSCA), flight trap; 20.IV.1982, Gordh, G., 1Ψ MEI 0 28378 (UCR); 12.IV.1983, Deyrup, M., 1ɗ MEI 162900 (FSCA), black light trap; 1.IV.1990, Wirth, W.W., 1ɗ MEI 0 79112, 10Ψ MEI 0 79110, 090356 –7, 090359 –64 (USNM), Malaise trap; 1.VI.1990, 1ɗ MEI 0 90358 (USNM), Malaise trap; 10.V.1983, Deyrup, M., 1Ψ MEI 162798 (FSCA), Malaise trap; 11.V.1983, 1Ψ MEI 162802 (FSCA), Malaise trap; 15.VII.1983, 1ɗ MEI 162901 (FSCA), Malaise trap; 25.I.1984, 1Ψ MEI 162797 (FSCA), Malaise trap; 11.VI.1984, 1Ψ MEI 162793 (FSCA), Malaise trap; 17.X.1984, 2Ψ MEI 162799, 162794 (FSCA), Malaise trap; 6.VI.1984, 1Ψ MEI 162792 (FSCA), Malaise trap; 25.X.1984, 1Ψ MEI 162796 (FSCA), Malaise trap; 29.XII.1984, 1Ψ MEI 162795 (FSCA), Malaise trap; 19.VII.1983, 1Ψ MEI 162811 (FSCA), Malaise trap; 23.VII.1983, 1Ψ MEI 162810 (FSCA), Malaise trap; 28.V.1984, 1ɗ MEI 162910 (FSCA), Malaise trap; 31.I.1986, 1Ψ MEI 162852 (FSCA), Malaise trap; 23.VII.1983, 1Ψ MEI 162809 (FSCA), Malaise trap; 3.VI.1985, 1ɗ MEI 162940 (FSCA), Malaise trap; Sebring [27.495, -81.441], 29.X.1955, Weems, Jr., H.V., 1ɗ MEI 112461 (USNM); 21.IV.1989, 1ɗ MEI 128532 (FSCA); 25.XII.1982, 1ɗ MEI 128480 (FSCA); 17.III.1975, 1ɗ MEI 162909, 2Ψ MEI 162239, 162276 (FSCA); 18.III.1975, 2ɗ MEI 162907–8, 8Ψ 162251, 162279, 162323, 162404, 162447, 162516, 162580, 162629 (FSCA), Malaise trap; 23.III.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 1ɗ MEI 162905 (FSCA), flight trap; 25.III.1978, 1ɗ MEI 162911 (FSCA), flight trap; 10.IV.1978, 1ɗ MEI 162879 (FSCA), flight trap; 17–18. IV.1978, 1 ɗ MEI 162903 (FSCA), flight trap; 23.IV.1978, 1ɗ MEI 162953 (FSCA), flight trap; 30.IV.1978, Weems, Jr., H.V., 1ɗ MEI 162915 (FSCA), flight trap; 31.IV.1978, Weems, Jr., H.V., Lampert, Jr., L.L., 1ɗ MEI 162913 (FSCA), flight trap; 1.VI.1978, Weems, Jr., H.V., Klein, L.K., 1ɗ MEI 162899 (FSCA), flight trap; 3.VI.1978, Weems, Jr., H.V., Klein, L.K., 1ɗ MEI 162919 (FSCA), flight trap; 30–31.VIII.1978, Weems, Jr., H.V., Lohrer, F.E., 1ɗ MEI 162906 (FSCA), flight trap; 14–18.IX.1978, Weems, Jr., H.V., Lohrer, F.E., 1ɗ MEI 162914 (FSCA), flight trap; 5.X.1978, Weems, Jr., H.V., Chance, S.J., 1ɗ MEI 162912 (FSCA), flight trap; 6.XII.1978, Weems, Jr., H.V., Chance, S.J., 1ɗ MEI 162918 (FSCA), flight trap; 30.XII–1.I.1979, Weems, Jr., H.V., Halkin, S., 1Ψ MEI 161167 (FSCA), flight trap; 23.II.1979, 1Ψ MEI 162551 (FSCA), flight trap; 26.III.1979, 1ɗ MEI 162931 (FSCA), flight trap; 31.V.1979, 1ɗ MEI 162929 (FSCA), flight trap; 4.VI.1979, 2Ψ MEI 161169 –70 (FSCA), flight trap; 5.VI.1979, 2Ψ MEI 161162, 162790 (FSCA), flight trap; 13–14.VII.1979, Weems, Jr., H.V., Webber, T.A., 1ɗ MEI 162936, 1Ψ 161168 (FSCA), flight trap; 23.VII.1979, 1ɗ MEI 162934 (FSCA), flight trap; 5.IX.1979, 1ɗ MEI 162935 (FSCA), flight trap; 7.IX.1979, 1ɗ MEI 162932 (FSCA), flight trap; 19.IX.1979, 1ɗ MEI 162933 (FSCA), flight trap; 3.X.1979, 2Ψ MEI 161164, 162756 (FSCA), flight trap; 4.X.1979, 1Ψ MEI 161166 (FSCA), flight trap; 5.X.1979, 2ɗ MEI 162930, 162937 (FSCA), flight trap; 9.X.1979, 2Ψ MEI 161171, 162754 (FSCA), flight trap; 19.X.1979, 1Ψ MEI 161165 (FSCA), flight trap; 10.IV.1964, Waters, J., 1Ψ MEI 118019 (FSCA); 14.IV.1968, Weems, Jr., H.V., 1Ψ MEI 162871 (FSCA), Malaise trap; 9.7 km W Sebring [27.495, -81.528], 15.III.1974, Smith, J.W., 1Ψ MEI 0 90174 (CMNH); W Sebring [27.471, -81.542], 17.IV.1989, Maier, C.T., 1ɗ MEI 111956 (CAES); 4.8 km S. FLA. 70 on US 27, 27.164, -81.328, 14.V.1964, Paulson, D.R., 1ɗ MEI 162920 (FSCA). Hillsborough County. 4.8 km S. FLA. 70 on US 27 [27.164, - 81.328], 14.V.1964, Paulson, D.R. 1ɗ MEI 162920 (FSCA); Plant City [28.02, -82.127], 22.III.1998, Hansen, D.E., 1ɗ MEI 133396 (MEIC); Tampa [27.947, -82.61], 2.II.1927, Bare, C.O., 1ɗ MEI 0 80354 (SEMC); 26.III.1961, Patton, J.V., 1Ψ MEI 0 0 3726 (FSCA); 21.II.1982, Eger, J. E., 4ɗ MEI 076474–5, 0 76477, 0 76479 (TAMU). Indian River County. Locality?, 24.III.1954, Mead, F.W., 1ɗ MEI 0 90385 (MEIC); Vero Beach [27.638, -80.398], Malloch, J.R., date?, 1ɗ MEI 112506, 3Ψ 112482, 112490 –1 (USNM). Lake County. Mount Dora [28.802, -81.645], 6.IV.1960, Felshaw, C.L., 1Ψ MEI 0 0 3709 (FSCA), fruit fly trap; 22.V.1962, Frederic, E.H., 1ɗ MEI 0 0 3668 (FSCA), Steiner trap; 24.V.1971, Castleman, D.L., 1ɗ MEI 162939 (UCR); 16.VII.1981, Jones, R., 1ɗ MEI 0 78309 (UMSP); Leesburg, W. & G. Lab. [28.811, -81.878], 8.IV.1963, Colon, J., 2ɗ MEI 128491, 128496 (FSCA); Fort Myers [26.64, -81.873], 24.IV.1930, Melander, A.L., 1ɗ MEI 112458 (USNM); 20.III.1980, Zeiss, M., 1ɗ MEI 0 75558 (CUIC), yellow pan. Lee County. Fort Myers [26.64, -81.873], 20.III.1980, Zeiss, M., 1ɗ MEI 0 75558 (CUIC), yellow pan. Liberty County. [30.569, -84.948], 13.VI.1974, Weems, Jr., H.V., Artaud, C.R., 1Ψ MEI 162867 (FSCA), Malaise trap. Manatee County. Palmetto [27.521, -82.573], 6.XII.1965, Chancey, D.C., 1Ψ MEI 0 0 3665 (FSCA), Steiner trap; Tallevast [27.401, -82.543], 17.V.1962, 1Ψ MEI 0 0 3720 (FSCA), fruit fly trap; 5.III.1930, Tinker, R.F., 1Ψ MEI 112485 (USNM); 25.IV.1961, Chancey, D.L., 1Ψ MEI 0 0 3704 (FSCA), fruit fly trap, 2.III.1962, 1ɗ MEI 0 0 3715 (FSCA), fruit fly trap; Oneco [27.447, -82.546], 31.I.1964, 1ɗ MEI 0 0 3677 (FSCA), Steiner trap; Piney Point [27.634, -82.541], 24.III.1972, Pierce, W.H., 1ɗ MEI 162952 (FSCA); 24.III.1972, Pierce, W.H., 1Ψ MEI 162869 (FSCA); Sarasota [27.336, -82.531], 23.III.1962, Yax, C.L., 1ɗ MEI 0 0 3716 (FSCA), Steiner trap; 30.IV.1963, 1Ψ MEI 0 0 3679 (FSCA), Steiner trap; 4.XI.1963, 1ɗ MEI 0 0 3675 (FSCA). Marion County. Ocala; [29.187, -82.14], 12.IV.1978, Wible, M., 1ɗ MEI 0 78445 (CMNH); Weirsdale [28.982, - 81.924], 19.IX.1963, Adkins, T.R., 1Ψ MEI 0 0 3650 (FSCA), Steiner trap; 29.IV.1964, 1Ψ MEI 0 0 3670 (FSCA), Steiner trap; Big Scrub [29.01, -81.8], 24–25.III.1979, Riley, E.G., LeDoux, 1ɗ MEI 0 39654 (UMRM). Miami-Dade County. Kendall [26.254, -80.124], 27.IX.1958, Butler, Jr., G.C., 1Ψ MEI 0 0 3661 (FSCA); Miami [25.774, -80.194], date?, Wight, A.E., 2ɗ MEI 162944, 162968 (MCZ); –.III.–, 1ɗ MEI 162967 (MCZ); 25.XI.1912, Knab, F.K., 1Ψ MEI 112474 (USNM); 19.XII.1912, 1ɗ MEI 112472 (USNM); 10.III.1932, Link, O.D., 1ɗ MEI 0 0 3671 (FSCA), Citrus paradisi ; 11.V.1932, Adams, B.F., 1ɗ MEI 0 0 3691 (FSCA), Bidens pilosa ; 11.II.1949, Calkins, O.W., 1ɗ MEI 0 0 3692 (FSCA), Eugenia uniflora ; 22.IV.1949, Link, O.D., 2Ψ MEI 003672–3 (FSCA); 9.III.1958, Dowling, Jr., C.F., 1ɗ MEI 0 0 3705 (FSCA); 6.III.1962, Harrisburg, 1Ψ MEI 162855 (MEIC); 20.II.1963, Brewton, W.S., 1ɗ MEI 0 0 3674 (FSCA); 1.V.1964, Paulson, D.R., 1Ψ MEI 162605 (FSCA); 21.III.1990, Rust, M.N., 1ɗ MEI 128494 (FSCA); Miami Springs [25.822, -80.29], 4.V.1965, Dowling, Jr., C.F., 1ɗ MEI 0 0 3680 (FSCA), McPhail trap. Orange County. Locality? 21.VII.1929, Hiers, W.A., 1Ψ MEI 112487 (USNM); –. X.1936, Link, O.D., 1ɗ MEI 112460 (USNM), McPhail trap; –. IV.1972, Wheeler, W.H., 1ɗ, UCF 0 55797 (UCFC); Conway [28.503, -81.331], 7.IX.1948, Link, O.D., 1Ψ MEI 0 0 3654 (FSCA); Maitland [28.618, -81.364], 6.VII.1979, 1Ψ, UCF 0 55799 (UCFC); Orlando [28.538, -81.379], –.II.–, collector?, 1ɗ MEI 162946 (MEIC); 28.II.1918, 1Ψ MEI 112479 (USNM); 4.III.1925, Cole, F.R., 1ɗ MEI 162966 (MEIC); 21.VII.1926, McBride, O.C., 1ɗ MEI 162965 (MEIC); –. III.1944, Bohart, R.M., Bohart, G.E., 2Ψ MEI 162856, 162858 (MEIC); 5.V.1977, Fullerton, S.M., 1Ψ, UCF 0 55804 (UCFC); 14.IV.1993, 1Ψ MEI 127214 (UCFC), Malaise trap, sand pine, turkey oak; Orlando, Florida Technological University (University of Central Florida) [28.6, -81.202], 20.VII.1972, 1ɗ, UCF 0 55802 (UCFC); 22.X.1990, Fullerton, S.M., 1Ψ, UCF 0 55801 (UCFC); 4.III.1991, 1Ψ, UCF 0 55763 (UCVC); 14.IX.1991, 1Ψ, UCF 0 55796 (UCFC). Orlando, VCF Arboretum [28.565, -80.705], 28.X.1993, Longhurst, J., 1Ψ, UCF 0 55810 (UCFC); US 50 and 436 [28.558, -81.313], 17.XI.1993, Anderson, M., 1ɗ, UCF 0 55808 (UCFC), sweep net, flatwoods; Winter Park [28.588, -81.351], –. III.1994, Johannsen, O.A., 1Ψ MEI 162860 (CUIC); 8.9 km N of Apopka [28.76, -81.51], 20.V.1978, Powers, J.R., 1Ψ MEI 0 82078 (MCZ). Pasco County. Blanton [28.411, -82.247], 7.VII.1965, Daniel, V.H., 1Ψ MEI 0 0 3652 (FSCA), Steiner trap; Dade City [28.364, -82.196], 20.X.1971, Williams, C.B., 1Ψ MEI 162851 (FSCA), McPhail trap; Elfers [28.217, -82.722], 3.VI.1929, Frierson, E.D., 2ɗ MEI 112480 –1 (USNM); Greenfield, 25.V.1967, Kennedy, O.D., 1ɗ MEI 0 0 3653 (FSCA), Steiner trap; Holiday [28.188, -82.74], 21.X.1983, Yackley, W.H., 1ɗ, CMN 55394 (CMNH); 17. II.1992, 1ɗ MEI 0 84374 (CMNH); 27. II.1992, 1ɗ MEI 0 84384 (CMNH); 29.V.1992, 1ɗ MEI 0 84385 (CMNH); 4.XI.1994, 1ɗ MEI 100196 (CMNH). Pinellas County. Locality?, 30.XI.1929, Moora, B.P., 1ɗ MEI 112493 (USNM); locality?, 29.VI.1953, Hill, L.B., 1Ψ MEI 0 0 3657 (FSCA); Dunedin [28.019, -82.772], –. I.1920, Blatchley, 1ɗ MEI 112459, 2Ψ MEI 112457, 112463 (USNM); 14.IV.1985, Dow, L.C., 1ɗ MEI 128533 (FSCA); 10.V.1985, 1ɗ MEI 128470 (FSCA); 12.V.1985, 1Ψ MEI 128475 (FSCA); 16.VI.1985, 1ɗ MEI 128535 (FSCA); Saint Petersburg [27.758, -82.642], 7.X.1963, Carroll, W.C., 1Ψ MEI 0 0 3678 (FSCA), Steiner trap; 6.XI.1964, Harris, S.B., 1ɗ MEI 0 0 3666 (FSCA), Steiner trap; 4.I.1974, Miller, R.C., 2ɗ MEI 0 75560, 0 75562 (CUIC); 21–26. XII.1982, 1 Ψ MEI 0 75559 (CUIC); Tarpon Springs [28.146, - 82.757], 18.X.1971, Williams, G.T., 1ɗ MEI 162960 (FSCA), McPhail trap; 18.X.1971, 1Ψ MEI 162868 (FSCA), McPhail trap. Polk County. Ft. Meade [27.752, -81.802], 25.II.1947, Burden, G.F., 1Ψ MEI 0 0 3729 (FSCA), at Citrus sinensis ; Lakeland [28.039, -81.95], 21.VIII.1962, Vild, R.E., 1ɗ MEI 0 0 3721 (FSCA), Steiner trap; Bartow [27.896, -81.843], –. V.1970, Pierce, W.H., 1ɗ MEI 0 28376 (FSCA), sweep net, water hyacinth. Putnam County. [Hollister], Hollister-Roberts Ranch [29.623, -81.814], 5.IV.1985, Milstrey, E.G., 1Ψ MEI 127471 (FSCA), gopher tortoise burrow; Palatka [29.648, -81.638], 28.IV.1957, Thompson, P.H., 1Ψ MEI 112453 (USNM). Sarasota County. Locality?, 3.III.1939, 1ɗ MEI 112495 (USNM); 5.IV.1961, Frederic, E.H., 1Ψ MEI 0 0 3710 (FSCA); Lido Key [27.317, -82.581], 18.IV.1975, Furth, D.G., 1ɗ, YPM 105212 (PMNH); Longboat Key [27.44, -82.69], 15.IV.1975, 1ɗ, YPM 105157, 1Ψ, YPM 105208 (PMNH); Sarasota [Memorial] Park [27.268, -82.527], 14.VI.1972, Pierce, W.H., 1ɗ MEI 162963 (FSCA). Seminole County.

[28.7, -81.2], 6.IV.1929, Dillers, Jr., B., 1ɗ MEI 112483 (USNM); 2.VI.1929, Sadler, J.E., 1Ψ MEI 112489 (USNM); 6.IX.1929, Hiers, Jr., B.D., 2ɗ MEI 112492, 112497 (USNM); 3.V.1950, Link, O.D., 1ɗ MEI 0 0 3656 (FSCA); 28.V.1971, Smith, W., 1ɗ, UCF 0 55798 (UCFC); 4.0 km N Astor Farms [28.865, -81.365], 18.IV.2000, Fullerton, S., Sims, T., 1Ψ, UCF 0 0 70127 (UCFC), Malaise trap, bald cypress swamp; Casselberry [28.678, -81.327], 15.VII.1959, Youtsey, C.O., 1ɗ MEI 0 0 3717 (FSCA); Longwood [28.694, -81.357], 11.VII.1979, Kaminski, R.L., 1Ψ, UCF 0 55800 (UFCC); Oviedo [28.673, -81.214], 28.V.1991, Fullerton, S.M., 1ɗ, UCF 0 55781 (UCFC), light trap; 13.II.1994, 1Ψ MEI 127212 (UCFC), Malaise trap; 21.II.1994, 1Ψ MEI 127211 (UCFC), Malaise trap; 11.III.1994, 1ɗ MEI 127695 (UCFC), Malaise trap; 22.III.1994, 1ɗ MEI 127698, 1Ψ MEI 127208 (UCFC), Malaise trap; 3.VII.1994, 1ɗ MEI 127699 (UCFC), Malaise trap; 10.VIII.1994, 1Ψ MEI 127697 (UCFC), Malaise trap; 20.VIII.1994, 2Ψ MEI 127696 127209 (UCFC), Malaise trap; 30.XI.1994, 1ɗ MEI 127213 (UCFC), Malaise trap; Paola [28.794, -81.363], 16.IX.1963, Desin, G.W., 1ɗ MEI 0 0 3676 (FSCA), Steiner trap; [28.81, -81.39], 15.VI.2001, Metz, M.A., MEI 130063 (INHS). St. John's County. North Fort Mantanzas, Anastasia Land [29.717, -81.238], 21.V.2002, Holston, K.C., 1ɗ MEI 140350 (INHS), coastal dunes; Saint Augustine Beach [29.85, -81.25], 20.IV.1969, Zeiger, C.F., 1Ψ MEI 162872 (FSCA); St. Augustine [29.894, -81.315], 1ɗ MEI 140538, 1Ψ 140539 (MCZ); – 15.V., collector?, 1Ψ MEI 0 80347 (SEMC); 15.III.1991, collector?, 1ɗ MEI 0 80348 (SEMC); 7.IV.1919, collector?, 1Ψ MEI 0 80897 (MEIC); 17.IV.1919, 1ɗ MEI 0 80895 (MCZ). St. Lucie County. Fort Pierce [27.446, -80.326], 13.VI.1962, Prance, E.E., 1Ψ MEI 0 0 3664 (FSCA), Steiner trap; 17.VIII.1981, Hibbard, K., 1Ψ MEI 128138 (FSCA), McPhail trap. Volusia County. Locality?, 27.VI.1929, Sadler, J.E., 1ɗ MEI 112488 (USNM); 12.VII.1981, Jones, R., 1ɗ MEI 0 78300 (UMSP), Virbirnum sp. 12.VII.1981, 1ɗ MEI 078303–7, 0 78311 (UMSP), Virbirnum sp. 13.VII.1981, 1ɗ MEI 0 78308, 0 78312 (UMSP), Virbirnum sp. 12.VII.1984, 3ɗ MEI 078301–2, 0 78310 (UMSP), Virbirnum sp. Daytona [Daytona Beach] [29.211, -81.023], – 8.IV., 1ɗ MEI 0 80899 (MCZ); 18.IV.1916, Johnson, C.W., 1ɗ MEI 162945 (MCZ); 6.IV.1919, 1ɗ MEI 162964 (MCZ); 8.IV.1919, 1Ψ MEI 0 80898 (MCZ); DeLand [29.028, -81.303], 29.VI.1951, Van Pelt, H.M., 1Ψ MEI 0 0 3655 (FSCA); 19.IV.1960, Desin, G.W., 1Ψ MEI 0 0 3711 (FSCA); 19.IV.1960, 1Ψ MEI 0 0 3712 (FSCA); Edgewater [28.989, -80.902], 24.II.1939, Frost, C.A., 1ɗ MEI 162951 (MCZ); Ormond Beach [29.286, -81.056], 22.VI.1939, Oman, 2ɗ MEI 112486 (USNM), 0 0 3743 (FSCA), 2Ψ 0 0 3742 (FSCA), 0 80896 (MCZ); Pierson [29.239, -81.466], 12.II.1962, Roberts, C.R., 1Ψ MEI 0 0 3725 (FSCA), Asparagus setaceus ; Blue Springs State Park [28.95, -81.34], 14.VI.1986, Powell, J., 1Ψ MEI 0 82091 (FSCA), blacklight. GEORGIA. Charlton County. Okefenokee Swamp, Billys Island [30.805, -82.341], –. VI.1912, 1 Ψ MEI 162859 (CUIC); –. VII.1912, 1 ɗ MEI 162955 (CUIC). Chatham County. Savannah [32.083, -81.1], 18.VI.1940, Hall, D.G., 1ɗ MEI 112496 (USNM). Liberty County. St. Catherines Island [31.657, -81.152], 24–28.IV.1972, Thompson, Picchi, 1ɗ MEI 0 76033 (SWRS). SOUTH CAROLINA. Paris Island, Beauford [32.436, -80.683], 2–8.X.1980, Reinert, J.F., 1Ψ MEI 128448 (FSCA). Horry County. Myrtle Beach [33.698, -78.895], 29.VI.1943, Parsons, C.T., 1Ψ MEI 162861 (MCZ). TEXAS. Cameron County. Locality?, 24.IV.1961, Eads, R.B., 1ɗ MEI 0 89875 (MEIC); Brownsville [25.901, -97.497], 29.VI.1938, Sailer, R.I., 1ɗ MEI 0 80353, 1Ψ MEI 0 80352 (SEMC); 3.VII.1938, Beamer, R. H., 1ɗ MEI 0 80350, 1Ψ 0 80351 (SEMC); 26.V.1939, Knull, D.J., Knull, J.N., 1Ψ MEI 112456 (USNM); 26.VI.1962, 1Ψ MEI 0 79726 (CNC); 10–13.III.1979, Friedlander, T., 1ɗ MEI 0 76476 (TAMU). Hidalgo County. Locality?, 19.X.1976, Fee, F.D., 2ɗ MEI 128485, 128488, 1Ψ 127287 (FSCA); [26.173, -98.382], 6.VI.1983, Barr, C.B., 1ɗ MEI 0 82178 (EMEC), blacklight; 15–16.III.1984, Carroll, L.E., Wharton, R.A., 1ɗ MEI 0 76478 (TAMU); 12 km ESE Hildalgo [26.065, -98.14], 25–26.III.1997, Webb, D.W., Metz, M.A., Holston, K.C., 1Ψ MEI 141427 (INHS), Malaise trap; 11.3 km S Alamo [26.067, -98.133], 25–27.VI.1980, Webb, D.W., Lisowski, E.A., 1Ψ MEI 141565 (INHS).

Figures

FIGURE 88. Penniverpa spp., distribution map. chersonesa (asterisk), epidema (open circle), festina (closed circle), insular (closed square), unispinosa (open square).

FIGURES 36 – 43. Penniverpa festina. 36. Tergite 8, epandrium, cerci, hypoproct dorsal view (inset ventral sclerite on hypoproct). 37. Sternite 8, hypandrium, gonocoxite ventral view. 38. Gonocoxite lateral view. 39. Gonostylus and ventral lobe dorsal view. 40. Aedeagus dorsal view. 41. Aedeagus ventral view. 42. Aedeagus lateral view. 43. Female internal reproductive organs. Scale = 0.1 mm. Abbreviations: AAP = accessory aedeagal process, C = cercus, CD = common duct, CSpD = common spermathecal duct, DA = dorsal apodeme, Dp = distiphallus, EA = ejaculatory apodeme, Epa = epandrium, F = furca, Gs = gonostylus, Gx = gonocoxite, H = hypandrium, Hpt = hypoproct, Igp = inner gonocoxal process, LGP = lateral gonostylar process, Sp = spermatheca, SpD = spermathecal duct, SpS = spermathecal sac, S 8 = sternite 8, T 8 = tergite 8, VA = ventral apodeme, VL = ventral lobe, VMP = ventromedial process.

FIGURES 78 – 87. Penniverpa spp., female abdomens. 78. alvatra, 79. chersonesa, 80. dives, 81. epidema, 82. evani, 83. festina, 84. gracilis, 85. insular, 86. megaplax, 87. multisetosa.