Helicops angulatus (Linnaeus, 1758)
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https://dx.doi.org/10.3897/BDJ.10.e69234 |
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persistent identifier |
https://treatment.plazi.org/id/D8B64A07-3420-537D-BA38-929FB9493E89 |
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treatment provided by |
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scientific name |
Helicops angulatus (Linnaeus, 1758) |
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Helicops angulatus (Linnaeus, 1758)
Materials
Type status: Other material. Occurrence: catalogNumber: MTKD 15294 ; recordedBy: Fritzsche leg.; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; Record Level: institutionID: MTKD Type status: Other material. Occurrence: catalogNumber: MTKD 15509 ; recordedBy: Poeppig leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: unknown province; locality: Ega ad Amazonas ; Event : year: 1831; Record Level : institutionID: MTKD Type status: Other material. Occurrence: catalogNumber: MTKD 41670 ; individualCount: 1; sex: female; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Peru; Record Level: institutionID: MTKD Type status: Other material. Occurrence: catalogNumber: SMF 100016 ; recordedBy: M. Jansen leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Bolivia; stateProvince: Santa Cruz; locality: Nuflo de Chavez , RPPN San Sebastian ; verbatimLocality: 524; verbatimLatitude: -16°23.263; verbatimLongitude: -61°59.983; Event : year: 2006; Record Level : institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 17817 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 17818 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 17819 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 17820 View Materials ; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 32409 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 32410 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 40029 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 80033 View Materials ; recordedBy: Edgar Lehr leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Peru; stateProvince: Ucayali; locality: Bolognesi (Campamento) ; verbatimLocality: 230; verbatimLatitude: -10°6.217; verbatimLongitude: -73°49.033; Event : year: 1998; Record Level : institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 90947 View Materials ; recordedBy: Gunther Köhler, R. Seipp, S. Moya leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Ecuador; stateProvince: Pastaza; locality: Arutam , km 48 Transamazonica ; verbatimLocality: 880; verbatimLatitude: -1°47.07; verbatimLongitude: -77°49.96; Event : year: 1996; Record Level : institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 91832 View Materials ; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 91833 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMF 91834 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMNS 13438 ; recordedBy: Hartmann leg.; individualCount: 1; sex: female; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Suriname; Event : year: 1893; Record Level: institutionID: SMF Type status: Other material. Occurrence: catalogNumber: SMNS 3063 ; recordedBy: F. Glocker leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Bahia; Event : year: 1854; Record Level : institutionID: SMNS Type status: Other material. Occurrence: catalogNumber: SMNS 3064.1; recordedBy: A. Kappler; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Event: year: 1843; Record Level: institutionID: SMNS Type status: Other material. Occurrence: catalogNumber: SMNS 3064.2; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: SMNS Type status: Other material. Occurrence: catalogNumber: SMNS 6394; recordedBy: A. Schlüter leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Event: year: 1985; Record Level: institutionID: SMNS Type status: Other material. Occurrence: catalogNumber: ZFMK 47670 ; individualCount: 1; sex: female; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Guyana; stateProvince: unknown province; locality: Roraima-Gebiet ; Record Level: institutionID: ZFMK Type status: Other material. Occurrence: catalogNumber: ZFMK 8403 ; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Amazonas; locality: Jurua ; Record Level : institutionID: ZFMK Type status: Other material. Occurrence: catalogNumber: ZFMK 8404 ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Amazonas; locality: Jurua ; Record Level : institutionID: ZFMK Type status: Other material. Occurrence: catalogNumber: ZMB 10854 View Materials ; recordedBy: S. Eye leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Maranhao; Record Level : institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 2303; recordedBy: M. Bloch leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 2305 ; individualCount: 1; sex: female; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: French Guiana; Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 25975 View Materials A ; recordedBy: K. Heller leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Suriname; stateProvince: Paramaribo; locality: Paramaribo ; Record Level : institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 25975 View Materials B ; recordedBy: K. Heller leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Suriname; stateProvince: Paramaribo; locality: Paramaribo ; Record Level : institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 26382 View Materials ; recordedBy: K. Heller leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Suriname; stateProvince: Paramaribo; locality: Paramaribo ; Record Level : institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 27783 View Materials ; recordedBy: Aq. Zoo. leg.; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 47771 View Materials ; recordedBy: K. Lako leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Para; locality: Rio Caramarapy ; Record Level : institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 54167 View Materials ; individualCount: 1; sex: female; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: French Guiana; Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 64697 View Materials ; recordedBy: Anat. Sammlung leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 64698 View Materials ; recordedBy: Anat. Sammlung leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 89648 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZMB 89649 View Materials ; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Record Level: institutionID: ZMB Type status: Other material. Occurrence: catalogNumber: ZSM 1525/0 ; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Suriname; Event : year: <1858; Record Level: institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 1526/0 ; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; Event : year: <1907; Record Level: institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 1528/0 ; recordedBy: Spix & Martius leg.; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; locality: in provinciae Bahiae adjacentibus ; Event : year: 1817-1820; Record Level: institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 247/1983 ; recordedBy: L. Müller leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus ( Linnaeus , 1758); Location : country: Brazil; stateProvince: Pará; locality: Peixeboi (an der Bragancabahn ) ; Event : year: 1910; Record Level : institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 264/2017 ; recordedBy: E. Snethlage leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Pará; locality: Rio Branco bei Obidos ; Event : year: 1912; Record Level : institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 37/2015 ; recordedBy: E.-G. Burmeister leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus ( Linnaeus , 1758); Location : country: Peru; stateProvince: Huánuco; locality: Biologische Station Panguana (unterer Rio Yuyapichis , ca. 140 km SSW Pucallpa) ; Event : year: 1982; Record Level : institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 516/2003 ; recordedBy: R. Zischka leg.; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Bolivia; stateProvince: Chapare ; Event : year: 1949; Record Level: institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 518/2003 ; recordedBy: H. Herkner leg.; individualCount: 1; sex: male; Taxon : scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Columbia; stateProvince: Guainía; locality: Inirida ; Event : year: 1976; Record Level: institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 59/1985 ; recordedBy: Maulhardt leg.; individualCount: 1; sex: female; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Peru; stateProvince: Ucayali; locality: Yarina Cocha bei Pucallpa ; Event : year: 1982; Record Level : institutionID: ZSM Type status: Other material. Occurrence: catalogNumber: ZSM 595/2003 ; recordedBy: O. Schindler leg.; individualCount: 1; sex: male; Taxon: scientificName: Helicops angulatus (Linnaeus, 1758); Location : country: Brazil; stateProvince: Sao Paulo; locality: Puerto Tiberica am Rio Parana [= Porto Tibirica] ; Event : year: 1938; Record Level : institutionID: ZSM
Diagnosis
Helicops angulatus can be distinguished from all its congeners, except H. scalaris and H. apiaka by having subcaudal keels, 17-20 dorsal scale rows at mid-body (compared to 19 in H. gomesi ; 21-27 in H. hagmanni ; 23-25 in H. pastazae ) and 103-123 ventrals (compared to 125-132 in H. gomesi ; 117-138 in H. hagmanni ). From H. scalaris , it differs in having no intergenials (for information on references, see Suppl. material 3; for summarised pholidosis information of the examined specimens, see Tables 1 View Table 1 , 2 View Table 2 ). For differentiation from H. apiaka , see identification of H. apiaka .
Distribution
The distribution of H. angulatus extends over nearly the complete northern part of South America. As shown in Fig. 1 a, the distribution range extends from Columbia to the east coast of Brazil and from Venezuela and offshore islands to the Brazilian Province of Sao Paulo and Bolivia.
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