Enicospilus tripartitus Chiu, 1954

Shimizu, So, 2020, The Nepalese species of the genus Enicospilus Stephens, 1835 (Hymenoptera, Ichneumonidae, Ophioninae): a preliminary revision and identification key to species, Deutsche Entomologische Zeitschrift 67 (1), pp. 69-126 : 69

publication ID	https://dx.doi.org/10.3897/dez.67.51332
publication LSID	lsid:zoobank.org:pub:2B601B5D-E1BD-44B7-BA89-554E3AB5EAE1
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treatment provided by	Deutsche Entomologische Zeitschrift by Pensoft
scientific name	Enicospilus tripartitus Chiu, 1954
status

Treatment

Enicospilus tripartitus Chiu, 1954 View in CoL Figs 2C View Figure 2 , 26 View Figure 26

Enicospilus tripartitus Chiu 1954: 36; holotype ♀, Taiwan, TARI, examined.

Material examined.

27♀♀ 10♂♂ and 2 unsexed: Nepal (24♀♀ 8♂♂ and 1unsexed), China (1♀), India (1♂), Japan (1 unsexed), Taiwan (2♀♀), unknown (1♂) .

Type series: holotype of Enicospilus tripartitus Chiu , 1954, ♀, Taihoku, Taiwan, 27.VIII.1937, J. Sonan leg. (TARI) ; paratype of same species, 1♂, no data (NHMUK).

Non-type series: 24♀♀ 8♂♂, Kakani (2,000 m), Nepal, VIII.1982 (2♀♀), VI (4♀♀ 1♂), VII (4♀♀ 3♂♂), 1-23 (2♀♀ 2♂♂).VIII (3♀♀ 1♂), IX (4♀♀), X (2♀♀). 1983, 1-30.V (1♀ 1♂), 1-14.VII (2♀♀).1984, M.G. Allen leg. (Figs 2C View Figure 2 , 26 View Figure 26 ); 1 unsexed, Sangu (ca 6,200′), Taplejung district , Nepal, 16-29.X.1961 ; 1♀, ShinKaiSi (1,340 m), Mt Omei , Szechuen, China ; 1♂, Kangra Valley (1,370 m), India, X.1899, Dudgeon leg. (all NHMUK); 1 unsexed, Genka-yama, Okinawa-hontô, Okinawa Pref., Japan, 4.V.1964, T. Takara & T. Kakinohana leg. (MNHA) ; 1♀, Kuanhsi , Taiwan, 29.VIII.1968 (MsT) (TARI) .

Distribution.

Eastern Palaearctic and Oriental regions ( Yu et al. 2016). Gauld and Mitchell (1981) recorded this species from Nepal.

Diagnosis.

Head (Figs 2C View Figure 2 , 26B-D View Figure 26 ): GOI = 2.2-2.9; lower face 0.7-0.8 × as wide as high; clypeus moderately to strongly convex in profile, its lower margin more or less blunt; mandible rather weakly twisted by 10-20°, moderately long, proximally tapered and distally parallel sided, its outer surface flat but with conspicuous dense setae and a proximal deep concavity; upper mandibular tooth 1.2-1.6 × as long as lower one; posterior ocellus close to eye; antenna with 55-66 flagellomeres and 20th flagellomere 2.2-2.4 × as long as wide.

Mesosoma (Fig. 26E View Figure 26 ): mesopleuron entirely more or less densely punctate and submatt; scutellum with lateral longitudinal carinae reaching posterior end and convergent posteriorly; metapleuron densely punctate as mesopleuron; propodeum weakly declivous, its posterior area moderately reticulate, outer margin of propodeal spiracle not joining pleural carina by a ridge.

Wings (Fig. 26F View Figure 26 ): fore wing with AI = 0.3-0.6, CI = 0.3-0.4, ICI = 0.5-0.7, SDI = 1.2-1.6; fore wing vein 1m-cu&M almost evenly curved to slightly sinuous, 2r&RS almost straight; fenestra and sclerites of discosubmarginal cell of fore wing as in Figure 26F View Figure 26 ; fenestra of fore wing not very long and its anterodistal corner distinctly separated from proximal end of vein RS; proximal sclerite triangular, separated from distal one, strongly pigmented; central sclerite strongly pigmented and sclerotised, well-delineated oval and its major axis parallel to distal margin of fenestra, positioned in mediodistal part of fenestra; distal sclerite absent to weak; proximal corner of marginal cell of fore wing uniformly setose; vein 1cu-a subinterstitial to antefurcal to M&RS by less than 0.2 × 1cu-a length.

Colour (Fig. 26 View Figure 26 ): body including interocellar area entirely reddish brown; wings hyaline.

Differential diagnosis.

Four Oriental Enicospilus species, E. laqueatus , E. pseudantennatus , E. vestigator , and E. tripartitus , have similar fenestra, sclerites, and fore wing veins (e.g. Figs 15F View Figure 15 , 21F View Figure 21 , 26F View Figure 26 ), as mentioned under E. laqueatus and E. pseudantennatus . Among them, E. tripartitus is readily distinguishable from other species by the outer mandibular surface: outer surface of mandible with very dense setae and sharp and rather deep proximal concavity in E. tripartitus (Figs 2C View Figure 2 , 26B, C View Figure 26 ), but more or less flat proximally with scattered setae in E. laqueatus (Fig. 15B, C View Figure 15 ) E. pseudantennatus (Fig. 21B, C View Figure 21 ) and E. vestigator .