Leiodidae, Fleming, 1821

Peck, Stewart B. & Cook, Joyce, 2014, A review of the small carrion beetles and the round fungus beetles of the West Indies (Coleoptera: Leiodidae), with descriptions of two new genera and 61 new species., Insecta Mundi 2014 (397), pp. 1-76 : 4-5

Key to subfamilies, tribes, and genera of Leiodidae of the West Indies

(Adapted in part from Peck et al. 1998)

1. Head with distinct occipital carina or crest. Cholevinae ............................................................. 2

— Head without occipital carina or crest. Leiodinae ....................................................................... 4

2(1). Mesocoxal cavities contiguous. Anemadini ............................................. Dissochaetus Reitter

— Mesocoxal cavities narrowly separated by prolonged mesosternal process ................................ 3

3(2). Head with strongly elevated occipital crest; maxillary palpomere 4 shorter than 3, conical; prosternum horizontal before coxae. Ptomaphagini ... Proptomaphaginus Szymczakowski

— Head with occipital crest weak or absent medially; maxillary palpomere 4 much longer than 3; prosternum flared ventrad to form precoxal cowling. Eucatopini .......... Eucatops Portevin

4(1). Labrum deeply emarginate apically; tarsal formula 5-5-4, not sexually dimorphic. Leiodini .. 5

— Labrum shallowly or not emarginate apically; tarsal formula 5-4-4 or less, or sexually dimorphic (5-5- 4 in male, less in female) .................................................................................................... 8

5(4). Antenna of 11 antennomeres with interrupted 5-antennomere club; antennomere 8 small and disc-shaped ......................................................................... Pseudolionothus Peck and Cook

— Antenna of 10 antennomeres with 3 or 4-antennomere club; antennomere 8 normal, rounded .. .................................................................................................................................................... 6

6(5). Mesotibiae broad; metatibiae slender, without strong spines on external margin ........................ ................................................................................................... Parvocyrtusa Peck and Cook

— Meso and metatibiae usually broad, shovel-like; metatibiae with strong spines on external margin ..................................................................................................................................................... 7

7(6). Antennal club of 3 antennomeres ..................................................................... Isoplastus Horn

— Antennal club of 4 antennomeres ................................................................ Zeadolopus Broun

8(4). Tarsal formula 3-3-3; first ventrite with transverse carina. Scotocryptini .............................. 10

— All tarsi 4- or 5-segmented; first ventrite without transverse carina. Agathidiini .................... 9

9(8). Tarsal formula 4-4- 4 in both sexes; parameres absent .............. Pseudoagathidium Angelini

— Tarsal formula 5-5- 4 in males, 5-4- 4 in females; parameres present ........ Agathidium Panzer