Ipomoea santillanii O'Donell

Wood, John R. I., Munoz-Rodriguez, Pablo, Williams, Bethany R. M. & Scotland, Robert W., 2020, A foundation monograph of Ipomoea (Convolvulaceae) in the New World, PhytoKeys 143, pp. 1-823 : 1

273. Ipomoea santillanii O'Donell , An. Inst. Biol. Mex. 12: 93. 1941. ( O’Donell 1941: 93)

Calonyction ventricosum Hallier f., Bot. Jahrb. Syst. 16: 556. 1894 [pub.1893]. (Hallier 1893a: 556), non Ipomoea ventricosa (Bertero) G. Don (1838) . Type. MEXICO. Veracruz, Valle de Cordoba, E. Bourgeau 1993 (holotype G00227297, isotype P).

Type.

Based on Caloncytion ventricosum Hallier f.

Description.

Perennial climber reaching 15 m, stems stout, glabrous, sometimes with occasional fleshy teeth. Leaves petiolate, very large, 15-20 × 12-18 cm, ovate, shortly acuminate, cordate, glabrous to thinly pubescent on the abaxial veins towards the base; petioles 10-20 cm. Inflorescence of long-pedunculate axillary cymes with up to 12 flowers, usually in a cymose cluster but sometimes (as in the type) forked into two branches, appearing lax and racemose; peduncles 6-30 cm; bracteoles caducous, suborbicular, convex, with fine terminal mucro c. 2-6 mm long, c. 2-3 cm × 1.5-2.5 cm, membranous with prominent venation; secondary peduncles up to 6.5 cm; pedicels 1-2.5 cm, thickened upwards; sepals similar, 9-14 × 4-7 mm, ovate-elliptic, obtuse, glabrous; corolla white, glabrous, funnel-shaped, basal cylindrical tube 1.5-5 cm long, 0.3-06 cm wide, then abruptly swollen before gradually being widened to mouth, the whole tube 4-6 cm long, the limb 2-3 cm long and 4-5 cm wide,; stamens shortly exserted. Capsules 2.5-3 × 1.5 cm, conical, rostrate, glabrous; seeds glabrous, 10 × 6 mm.

Illustration.

Figure 8F .

Distribution.

Southern Mexico to Costa Rica. Disturbed woodland on stony ground, 800-1300 m.

COSTA RICA. San José, P. Döbbeler 934 (BM); Río María, A. Tonduz 8439 (BM), 13051 (K).

EL SALVADOR. La Libertas, Antigua Cuscatlan, P. Lemus s.n. [13/1/1989] (K, LAGU).

GUATEMALA. Chapadero, Santa Rosa, Heyde & Lux 4354 (K).

MEXICO. Chiapas: Mun. San Fernando, J. Carmen Soto et al. 13324 (BM, MEXU, ARIZ); Esquintla, E. Matuda 17292 (K). Colima: Rancho El Jabali, E. Lott et al. 3009 (MICH). Est. México & Dist. Fed.: Temascaltepec, Tejupilco, G.B. Hinton 2296 (BM, K); ibid., Luvianos, G.B. Hinton 7169 (K); Piedras Negras, E. Matuda 29702 (IEB). Guerrero: El Balsamo, J.C. Soto Nuñez 11469 (MEXU). Jalisco: San Sebastián del Oeste, T.S. Cochrane et al. 12045 (IEB). Michoacán: Coalcomán, Sierra Naranjillo, G.B. Hinton 12680 (K); Zirimicuaro, S. Zamudio 11264 (IEB). Oaxaca: Santa Cruz Tepitotula, P. Osorio 265 (IEB). Veracruz: Orizaba, E. Bourgeau 3024 (P, K); Moyapan, Orizaba, M. Rosas 709 (BM. GH, MEXU).

Note.

The variations in inflorescence structure and corolla size are so great that it is difficult to believe only one species is involved.

Figures

Figure 8. Corollas showing variations in form (side view), size, limb lobing and stamen exsertion. A Ipomoea argentea B I. repanda C I. neei D I. electrina E I. habeliana F I. santillanii G I. nationis H I. rubriflora J I. longistaminea K I. megapotamica L I. megalantha M I. neriifolia N I. syringifolia P I. ramosissima Q I. elongata R I. mucronatoproducta. A from Wood et al. 25639 and photo; B from Whitefoord 5244; C from Skutch 2043; D from Breedlove 27626; E from Bentley 203; F from Bourgeau 3024; G from Saunders 987; H from Wood et al. 27678; J from Pastore et al. 2678; K from Wood et al. 28060; L from Hassler 9114; M from Rezende et al. 1011; N from Stutz 1426 and photo; P from Bang 2246; Q from Purpus 3904; R from Wood & Villarroel 25474. Drawn by Rosemary Wise.