Pusia moravica ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 41

Pusia moravica ( Hoernes & Auinger, 1880) nov. comb.

Figs 13I 1 –I 2 , J

* Mitra plicatula Var. ( moravica )— Hoernes & Auinger 1880: 84.

Type material. Lectotype (designated herein): NHMW 1862 /0029/0006, SL: 9.8 mm, MD: 4.7 mm, Rudice ( Czech Republic), figs 13I 1 –I 2 . Paralectotype: NHMW 2020 /0097/0001, SL: 12.6 mm, MD: 6.5 mm, Rudice ( Czech Republic), fig. 13J .

Revised description. Shell moderately large, stout fusiform, with slightly gradate spire. Protoconch and early teleoconch unknown. Teleoconch whorls low, weakly convex, with periphery in abapical third; suture impressed. Sculpture of sharp crested, orthocline to weakly prosocline ribs, separated by interspaces of about same width (about 15 axial ribs on penultimate whorl). Adapical weak subsutural spiral groove cuts ribs, forming narrow subsutural spiral cord of weak nodes. Further delicate spirals placed below subsutural cord, abapically rapidly weakening. Last whorl convex with short, moderately constricted base. Axial ribs becoming subobsolete on last whorl. Columella with four prominent, oblique columellar folds. Siphonal canal moderately long, straight.

Shell measurements and ratios. SL: 9.8–12.6 mm, MD: 4.7–6.5 mm (fragments; restored height: 13–16 mm); assumed SL of complete specimen: 18 mm, AA: 33°.

Discussion. Only two fragments are available, which Hoernes & Auinger (1880) described as variety of “ Mitra plicatula ” (= Vexillum transalpinum nov. sp.). A close relationship between Pusia moravica ( Hoernes & Auinger, 1880) and V. transalpinum , however, is unlikely in respect to the short spire whorls and broad, convex last whorl of P. moravica . In addition, the subsutural spiral row of weak nodes and the presence of spiral grooves allows a clear separation from V. transalpinum .

Palaeoenvironment. The specimens derive from basinal clay, suggesting a middle neritic depositional environment.

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Rudice ( Czech Republic) ( Hoernes & Auinger 1880).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 13 A1–A2. Pusia avellanella (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360343. B1–B2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0002. C1–C2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. D1–D2. Pusia avellanella (Boettger, 1906), Coşteiu de Sus (Romania), SMF 360344. E1–E2. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0004. F1–F2. Pusia avellanella (Boettger, 1906), Steinebrunn (Austria), NHMW 1847/0037/0048. G1–G2. Pusia falsitranssylvanica nov. nom., holotype, Coşteiu de Sus (Roma- nia), SMF 360346. H1–H2. Pusia confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. I1–I2. Pusia moravica (Hoernes & Auinger, 1880), lectotype, Rudice (Czech Republic), NHMW 1862/0029/0006. J. Pusia moravica (Hoernes & Auinger, 1880), paralectotype, Rudice (Czech Republic), NHMW 2020/0097/0001.