Pusia pseudorecticosta ( Boettger, 1906 )

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 42-44

Pusia pseudorecticosta ( Boettger, 1906)

Figs 4J , 15A 1 –A 2 , B 1 –B 2 , C 1 –C 2 , D

Mitra (Costellaria) recticosta Bell. — Hoernes & Auinger 1880: 85, pl. 10, figs 5–8 [non Bellardi, 1850].

Mitra (Costellaria) recticosta Bell. — Mikhailovsky 1903: 202.

* Mitra (Costellaria) pseudorecticosta n. nom.— Boettger 1906: 10.

Mitra dubia n. sp. — Friedberg 1910: 104 [non Hutton, 1873].

Turricula vindobonensis Friedb. — Friedberg 1911: 26, pl. 1, fig. 18.

Turricula (Uromitra) vindobonensis Friedb. — Meznerics 1933: 343, pl. 14, figs 7a–b.

Vexillum (Costellaria) recticostatum (Bell.) — Sieber 1956: 244.

V [exillum]. (C [ostellaria].) vindobonense (Fbg.) — Sieber 1958a: 153.

Vexillum (Costellaria) vindobonense (Fbg.) — Sieber 1958b: 149.

Vexillum (Costellaria) pseudorecticosta (Boettger, 1901) — Bałuk 1997: 36, pl. 10, figs 3–7.

Pusia transsylvanica ( Boettger, 1902) — Landau et al. 2013: 216, pl. 34, figs 10–12 [non Boettger, 1902].

non Vexillum (Costellaria) vindobonense Friedberg — Csepreghy-Meznerics 1954: 45, pl. 5, fig. 30 [=?].

non Vexillum (Costellaria) cfr. vindobonense (Friedberg, 1923) — Švagrovský 1981: 51, pl. 45, fig. 8 [=?].

Type material. Lectotype (designated herein): NHMW 2020 /0076/0002, SL: 10.2 mm, MD: 3.6 mm, Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1880: pl. 10, figs 6a–c), figs 15A 1 –A 2 . Paralectotype: NHMW 2020 /0076/0001, SL: 16.0 mm, MD: 5.5 mm, Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1880, pl. 10, figs 5a–c), figs 15B 1 –B 2 .

Additional material. NHMW 2020 /0076/0003, SL: 14.4 mm, MD: 4.9 mm, Steinebrunn ( Austria) , fig. 15D; NHMW 2020 /0076/0004, SL: 14.0 mm, MD: 5.0 mm, Steinebrunn ( Austria) , figs 15C 1 –C 2; NHMW 1887 /0018/0019, Steinebrunn ( Austria) , illustrated in Meznerics (1933, pl. 14, figs 7a–b); NHMW 2020 /0076/0005, SL: 9.8 mm, MD: 3.7 mm, Steinebrunn ( Austria) , fig. 4J; NHMW 1855 /0045/0035, 80 specimens, Steinebrunn ( Austria) ; NHMW 1866 /0001/1018, 24 specimens Forchtenau ( Austria) ; NHMW 1871 /0010/0015, 2 specimens, Sedlec (= Porzteich) ( Czech Republic) .

Revised description. Shell moderately large, moderately slender fusiform, with high, slightly gradate spire. Protoconch ( Fig. 4J ) multispiral, high conical of 3.5 smooth, nearly straight-sided whorls (height: 940 μm, diameter: 750 μm). Teleoconch of about eight subcylindrical to weakly convex whorls, separated by narrowly impressed suture. Axial sculpture of close-set orthocline ribs separated by slightly narrower interspaces (about on 20 penultimate whorl). Last whorl strongly constricted. Axial sculpture on last whorl often reduced. Aperture wide, ovoid with indistinct anal canal. Columellar callus indistinct, barely delimited from base. Columella with four oblique columellar folds, weakening abapically. Outer lip convex, thin, with 10–12 long delicate lirae placed deep within aperture. About five weak to prominent spiral cords on weak fasciole, partly extending from columellar folds. Siphonal canal moderately short, bent slightly to left, with shallow siphonal notch.

Shell measurements and ratios. SL: 10.2–16.0 mm, MD: 3.6–5.0 mm; AA: 32–40°, SL/MD: 2.8–2.9, AL/AW: 3.0–3.9, AH/S: 2.5–2.7.

Discussion. Hoernes & Auinger (1880) identified this species as Mitra recticosta , which was described by Bellardi (1850) from the Pliocene of Italy. Bellardi (1887b) doubted this identification and separated the Viennese shells based on their smaller size, more numerous and narrower axial ribs and the shorter last whorl from his M. recticosta but did not propose a new name. This was done by Boettger (1906) who introduced Mitra pseudorecticosta as new name. It is important to note that although Boettger (1906) described specimens from Coşteiu de Sus (Romania), he clearly referred to the Austrian specimens illustrated by Hoernes & Auinger (1880), when introducing Mitra pseudorecticosta as new name. Curiously, the specimens from Coşteiu de Sus, which Boettger (1906) had at hand when discussing Mitra pseudorecticosta are not conspecific with the Austrian shells and are described here as Pusia confunda nov. sp. Friedberg (1910), who was unaware of Boettger’s paper, introduced Mitra dubia as new name for Mitra recticosta Hoernes & Auinger, 1880 . This name, however, was preoccupied by Hutton (1873) and therefore, Friedberg (1911) proposed Turricula vindobonensis as replacement name a year later. Boettger (1906) and Friedberg (1910, 1911) refer to the same lot of illustrated specimens and therefore, Turricula vindobonensis is an objective junior synonym of Mitra pseudorecticosta Boettger, 1906 . The placement of this species in Pusia is tentative.

Hölzl (1973: 93) mentioned Vexillum (Costellaria) vindobonense (Friedberg) from the Ottnangian (middle Burdigalian) of Kaltenbachgraben in Bavaria. This would be the oldest record of this group from the Paratethys Sea.

Palaeoenvironment. Most abundant at localities suggesting silty-sandy inner neritic environments partly with sea grass meadows (e.g. Steinebrunn) (own data).

Distribution in Central Paratethys. Badenian (middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1997); Ukrainian Fore-Carpathian Basin: Dryszczów (Nadrichne), Zborów (Zboriv) ( Ukraine) ( Friedberg 1911); Vienna Basin: Niederleis, Steinebrunn ( Hoernes & Auinger 1880), Sedlec (= Porzteich) ( Czech Republic); Eisenstadt-Sopron Basin: Forchtenau ( Hoernes & Auinger 1880).

Eastern Paratethys. Tarkhanian (early Middle Miocene) of Tomakivka (S. Ukraine) ( Mikhailovsky 1903). Proto-Mediterranean Sea. Karaman Basin: Seyithasan ( Turkey) ( Landau et al. 2013).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 4. Protoconchs of Paratethyan Costellariidae. A. Antithala filipescui nov. sp., Lăpugiu de Sus (Romania), NHMW 1876/0011/0012). B. Antithala sturi (Hoernes & Auinger, 1880), Lăpugiu de Sus (Romania), NHMW 2020/0058/0001. C. Bellardithala kovaci nov. sp., Steinebrunn (Austria), NHMW 2020/0062/0001. D. Bellardithala partschi (Hörnes, 1852), Steinebrunn (Austria), NHMW 2020/0060/0003. E. Ebenomitra leucozona (Andrzejowski, 1830), Lăpugiu de Sus (Romania), NHMW 1876/0011/0008. F. Ebenomitra pseudopyramidella (Boettger, 1906), Steinebrunn (Austria), NHMW 1857/0014/0009. G. Fedosovia fuchsi (Hoernes & Auinger, 1880), lectotype, Lăpugiu de Sus (Romania), NHMW 1857/0024/0012. H. Pusia avellanella (Boettger, 1906), Gainfarn (Austria), NHMW 2020/0072/0001. I. Pusia? confunda nov. sp., holotype, Coşteiu de Sus (Romania), SMF 360341. J. Pusia pseudorecticosta (Boettger, 1906), Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0005. K. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. L. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. M. Vexillum intermittens (Hoernes & Auinger, 1880), paralectotype, Baden or Bad Vöslau (Austria), NHMW 2020/0087/0003. N. Vexillum michelottii (Hörnes, 1852), lectotype, Baden (Austria), NHMW 1851/0013/0001. O. Tosapusia pseudocupressina Bałuk, 1997, Lăpugiu de Sus (Romania), NHMW 1870/0033/0037.

FIGURE 15 A1–A2. Pusia pseudorecticosta (Boettger, 1906), lectotype, Steinebrunn (Austria), NHMW 2020/0076/0002. B1–B2. Pusia pseudorecticosta (Boettger, 1906), paralectotype, Steinebrunn (Austria), NHMW 2020/0076/0001. C1–C2. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0004. D. Pusia pseudorecticosta (Boettger, 1906), Steinebrunn (Austria), NHMW 2020/0076/0003. E1–E2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0002. F1–F2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0003. G1–G2. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0078/0001. H. Pusia schafferi (Meznerics, 1933), Steinebrunn (Austria), NHMW 2020/0079/0001. I1–I2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2020/0081/0001. J1–J2. Pusia vexans (Boettger, 1902), lectotype, Coşteiu de Sus (Romania), SMF 360339. K1–K2. Pusia vexans (Boettger, 1902), Coşteiu de Sus (Romania), NHMW 2018/0248/0327.