Viidalepp, Jaan, 2011, A morphological review of tribes in Larentiinae (Lepidoptera: Geometridae), Zootaxa 3136, pp. 1-44: 19

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http://doi.org/ 10.5281/zenodo.279481



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Tribe Rheumapterini   Herbulot, 1961

( Figs 62–63, 65 – 71 View FIGURES 62 – 68 View FIGURES 69 – 73 )

Herbulot (1961) separated the tribe without a diagnosis. Large dark-coloured moths with distally dentate-edged hind wings have always been kept together in check-lists. However, there are three distinct groups of genera: the Rheumaptera   informal group of related genera, the large-sized troglophilous species of Triphosini, both are treated below as tribes Rheumapterini   and Triphosini, respectively. The third, Phileremini consists of a few Palaearctic species with several specific traitsas listed below.

The tribe Rheumapterini   shows a long list of apomorphic characters: juxta dorsal part rooted in the basal part of it ( Figs 62–65 View FIGURES 62 – 68 ); a slender sclerite present between juxta, saccus and bases of sacculi ( Fig. 62 View FIGURES 62 – 68 ). It is not clear whether this sclerite is analogous to that in Cataclysmini; The rheumapterine genera are characterized here by: wide posterior sclerotisation of female genitalia (possibly shared with Melanthiini   ?) ( Figs 66–68 View FIGURES 62 – 68 ); uncus heavily sclerotized, broad, flat, triangular, trapezoidal or cupola- shaped (skinny soft and hairy in Melanthiini   ); both costal and saccular ornamentations of valva present (like also in Melanthiini   ); labides arise from the base of costa, are connected by a membrane, supported by a median sclerite, long, headed, reaching uncus ( Figs 62–65 View FIGURES 62 – 68 ); dorsal parts of vinculum flattened and invaginated into bases of valvae; presence of coremata associated with membranization of the eighth abdominal segment in male and large masses of deciduous hair-scales ( Fig. 71 View FIGURES 69 – 73 ).

Some apomorphic characters of subclades occur within the tribe, such as: shortening of spurs on male hind tibia; rich vestiture of male hind tibia associated with shortened and thickened tibia, and the basal tarsomere ( Figs 69, 70 View FIGURES 69 – 73 ); the presence of blotches of specialized scales in wings of male; posterior sclerotization of female genitalia; fusion of cornuti in male aedeagus and signa in female bursa copulatrix into conglomerates.

Distribution: Neotropical, Holarctic.