Cyclocephala gravis Bates, 1888

Moore, Matthew R., Cave, Ronald D. & Branham, Marc A., 2018, Annotated catalog and bibliography of the cyclocephaline scarab beetles (Coleoptera, Scarabaeidae, Dynastinae, Cyclocephalini), ZooKeys 745, pp. 101-378 : 168

Cyclocephala gravis Bates, 1888

Cyclocephala gravis Bates, 1888: 308 [original combination].

syn. Cyclocephala meinanderi Endrődi, 1964: 457-459 [original combination].

Cyclocephala gravis ab. meinanderi Endrődi [new infrasubspecific status by Endrődi 1967: 90].

Cyclocephala gravis Bates [synonymy by Maes 1994: 11].

Types.

Type of C. gravis at BMNH ( Endrődi 1966). Holotype ♂ of C. meinanderi at HNHM ( Endrődi Collection) ( Endrődi 1964).

Distribution.

BELIZE: Cayo, Orange Walk, Stann Creek, Toledo. BOLIVIA: Cochabamba, Santa Cruz. BRAZIL: Bahia, Espírito Santo, Mato Grosso, Minas Gerais, Pernambuco, Rio de Janeiro. COLOMBIA: Antioquia, Boyacá, Caquetá, Chocó, Cundinamarca, Magdalena, Santander, Tolima. COSTA RICA: Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas, San José. ECUADOR: Santo Domingo de los Tsáchilas. EL SALVADOR: Morazán. FRENCH GUIANA: Cayenne, St.- Élie, St.-Laurent du Maroni. GUATEMALA: Alta Verapaz, Huehuetenango, Izabal, Petén, Sololá, Zacapa. GUYANA: Upper Demerara-Berbice. HONDURAS: Atlántida, Choluteca, Cortés, El Paraíso, Francisco Morazán, Gracias a Dios, Olancho, Santa Bárbara, Yoro. MEXICO: Campeche, Chiapas, Guanajuato, Oaxaca, Quintana Roo, San Luis Potosí, Tabasco, Veracruz, Yucatán. NICARAGUA: Chontales, Jinotega, Matagalpa, RAA Norte, Río San Juan. PANAMA: Bocas del Toro, Chiriquí, Colón, Former Canal Zone, Panamá. PARAGUAY: Distrito Capital. SURINAME. VENEZUELA: Capital District, Guárico.

References.

Bates 1888, Bodkin 1919, Arrow 1937b, Blackwelder 1944, Pike et al. 1976, Dechambre 1979a, 1980, Endrődi 1964, 1966, 1967, 1985a, Young 1986, 1988a, b, 1990, Thomas 1993, Lobo and Morón 1993, Maes 1987, 1994, Ramírez and Brito 1992, Croat 1997, Ratcliffe and Morón 1997, Beath 1998, 1999, Ratcliffe 1992b, 2002a, 2003, Restrepo et al. 2003, Reyes Novelo and Morón 2005, Ratcliffe and Cave 2006, Krajcik 2005, 2012, Neita-Moreno 2011, Ponchel 2011, García- López et al. 2013, Moore and Jameson 2013, Ratcliffe et al. 2013, López-García et al. 2015, Gasca-Álvarez and Deloya 2016.