Libellula Linnaeus 1758

Mauffray, William F. & Tennessen, Kenneth J., 2019, A Catalogue and Historical Study of the Odonata of Ecuador, Zootaxa 4628 (1), pp. 1-265 : 171-176

Libellula Linnaeus 1758

Of the 28 described New World species, only one species has been found in Ecuador ( Fig. 99 ), the nymphs of which develop in small shaded pools and ditches.

Libellula herculea Karsch, 1889 . Publ. Recs. ECU: Kennedy (1922), Racenis (1954), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); AZ: * Kirby (1889 —as Belonia longipennis ), Calvert (1906), Campos (1922); BO: Ris (1910); ES: Calvert (1909), Campos (1922), Daigle et al. (1997); OR: von Ellenrieder & Garrison (2011); PC: * Calvert (1909), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, VEN: (H06), (P11); plus SUR: (P16).

Addl. Data. AZ, Santa Isabel: 1 ♂, stream, 21 km E of Hwy. 25, road from Balao jct. (-2.9400, -79.5653, 730m), 11 iv 2008, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. JTJ; BO, San Miguel: 1 ♂, Bal- zapamba (-1.7667, -79.1833, 700m), ix 1935, leg. F. Campos, det. MJW [ANSP]; EO, Piñas: Buenaventura, Río Principal (-3.6561, -79.7735, 329m), 19 vi 2016, leg. & det. ACRP [ACRP]; ES, San Lorenzo: Lita along RR track, 1–5 km W, adjacent to Río Baboso seeps, pools & streams (0.8792, -78.4589, 535m), 5 ii 1997, 5 ♂♂, 1 ♀, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 1 ♂, 1 ♀, same data but leg. & det. JJD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; MS, Huanboya : 1 ♂, small stream near Río Pasanac , Hwy . 45, S of river, E side of Hwy . , 16 ix 2005, leg. & det. KJT [KJT]; 1 ♀, Macas , 58 km N on Hwy . 45 (70 km N of Sucua), roadside ditch and tribs near Río Pasanac (-1.9447, -77.8536, 762m), 10 xi 1997, leg. & det. WFM [FSCA]; Taisha : 1 ♀, Macuma , small ponds (867m), 1 ii 1972, leg. D. Pearson, det. DRP [DRP]; NA, Tena: 2 ♂♂, Tena (500m), leg. F. Campos, det. MJW [ANSP]; 1 ♂, Tena (-0.98, -77.80, 500m), date?, leg. F. Campos, det. MJW [FSCA]; adult, 1 km S of Tena , 28 v 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, J. S.B. S., near Tena (450m), 18 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♂, T. B.S., open grassy area behind dining room, surrounded by bushes and trees and some seepage areas (-0.6383, -76.1492, 220m), 10 i 2009, leg. & det. RWG & NVE [USFQ]; Loreto: 1 ♀, small streams and forest trails 12 km E of Loreto, 2+ km S of Hwy. 45 (-0.6217, -77.2600, 365m), 12 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, Cotapino, S along roadside stream (-0.7076, -77.4743, 530m), 4 xi 2013, leg. & det. CDK [FSCA]; same data but 1 ♂, leg. & det. KJT [FSCA]; PC, San Miguel Bancos: 1 ♂, Puerto Quito (750m), 28 v, leg.?, det. SWD [QZAC]; 1 ♂, 1 ♀, Mindo, swamps and streams near town (-0.0556, -78.7714, 1244m), 28 i 2009, leg. & det. RWG & NVE [RWG]; 1 ♂, 1 ♀, Mindo, Via al Mariposario, Mindo Gardens Eco Lodge, vege- tated cement pond (-0.0720, -78.7560, 1390m), 3 ix 2018, leg. & det. KJT [KJT]; Quito : 3 ♂♂, 1 ♀, Río Tulambi , Forestry Sta. Maquipucuna , about 5 km E of Nanegal (0.12, -78.62), 9 v 1990, leg. OSF, det.? [USNM]; SD: 1 ♀, Santo Domingo, 12 km E at Tinalandia , 11–17 v 1986, leg. G. Edwards, det. SWD [SWD]; 2 ♂♂, about 93 km SW Quito, Chiriboga Road , 9 vi 1990, leg. OSF, det.? [USNM]; SU, Shushufindi: 1 ♀, Limoncocha , forest trail, 8 x 1971, leg. D. Pearson, det. DRP [DRP].

Note: There is a ♂ specimen in QZAC labeled “Calvla” 12 i 1977; we could not find this location in any reference.

Macrothemis Hagen, 1868

A species-rich genus of 41 species with at least 14 species in Ecuador. The latest key to males was provided by May (1998) although species identification is difficult; females of very few species are identifiable. These are small to medium-sized mostly dark dragonflies ( Fig. 100 ) that inhabit small streams to medium-sized rivers, both open and forested.

Macrothemis delia Ris, 1913 . Publ. Recs. E CU: *(H14). Also CEN, SUR, VEN: (H06), (P16).

Addl. Data. OR, Francisco de Orellana: 1 ♂, Río Shiripuno , bridge crossing 59 km SSE of Coca, (-1.0079, -76.9451), date not provided, data provided by WAH & DLW; 1 ♂, T. B.S., Río Tiputini , 55 km SE of Pompeya, 94 km ESE of Coca (-0.6369, -76.1503), date not provided, data provided by WAH & DLW.

* Macrothemis extensa Ris, 1913 . Publ. Recs. BRA, CEN, GUF, PER, VEN: (H06), (P11); plus SUR: (P16).

Addl. Data. LR, Buena Fé: 1 ♂, Río Palenque Biological Station, 47 km S of Santo Domingo de los Colorados (0.5864, -79.3647, 150m), 8 x 1988, leg. & det. SWD [SWD]; SD: 1 ♀, Alluriquín 7.3 km W of Hotel Tinalandia, 29 x 1994, leg. A. Jaramillo, det. DRP [DRP].

* Macrothemis fallax May, 1998 . Publ. Recs. CEN: May (1998), (P16); plus COL: (P&P), (P16).

Addl. Data. ES, San Lorenzo: 1 ♀, tiny stream near RR track, about 4 km N of Lita, 5 ii 1997, leg. & det. KJT [KJT]; MN, Portoviejo : 1 ♂, concrete ditch, Hwy. E 40, E of Portoviejo, 4 iii 2010, leg. & det. JJD [JJD]; 1 ♂, same data but leg. KJT, det. WFM [FSCA]; SD: 3 ♂♂, Tinalandia , 12 km E of Santo Domingo de los Colorados, streams S of river (-0.2839, -79.05, 760m), 5 x 1988, leg. & det. SWD [RWG, SWD].

Note: The additional records are the southernmost for this species which is often confused with M. pseudimitans .

Macrothemis hahneli Ris, 1913 . Publ. Recs. ECU: (H06), (P11), (H14). * Daigle et al. (1996). NA: Daigle (1996— misidentified as M. musiva ). Also in COL, PER, VEN: (H06), (P11); plus ARG: (Garrison pers. comm. 2019).

Addl. Data. MS, Huamboya: 1 ♂, Macas, 58 km N on Hwy. 45 [70 km N of Sucua], roadside ditch and tribs near Río Pasanac (-1.9447, -77.8536, 762m), 16 ix 2005, leg. & det. WFM [FSCA]; Morona: 3 ♂♂, 1 ♀, small stream and roadside seepage pool, 15 km W of Macas (-2.2731, -78.1939, 1070m), 15 ix 2005, leg. & det. KJT [KJT]; 3 ♂♂, 1 ♀, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, rocky stream, 22.5 km W of Macas, Guamote Road, 18 ix 2005, leg. & det. JJD [JJD]; 2 ♂♂, 2 ♀♀, Macas, 22.9 km W on Guamote Road, 3.3 km W on Río Al- banico rocky stream and flowing roadside ditches (-2.2342, -78.2181, 1690m), 18 xi 2005, leg. & det. WFM [FSCA, LSAM]; NA, Archidona: 5 ♂♂, 1 ♀, [Narupa], E on road to Loreto, 43.5 km W of Loreto seep near large rocky river [Río Hollin] (-0.7022, -77.7189, 1036m), 15 vii 1996, leg, & det. JJD [JJD]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 14 vi 1995, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but 15 vii 1996, leg. & det. WFM [FSCA, LSAM]; 5 ♂♂, same data but 14 xi 1997 [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, [Sarayacu], 12.8 km N of Narupa on Baeza Hwy., small shallow sandy stream (-0.67, -77.78, 1585m), 14 vii 1996, leg. & det. WFM [LSAM, QZAC]; 3 ♂♂, same data but leg. KJT [KJT]; 3 ♂♂, same data but leg. & det. SWD [SWD]; 2 ♂♂, seep by waterfall, Loreto Road, 44.1 km E of Narupa jct., 14 xi 1997, leg. & det. KJT [KJT]; 1 male same data but leg. & det. TWD [SWD]; 2 ♂♂, [Jondachi], 42 km N of Tena on road between Tena & Baeza (-0.72, -77.78, 1190m), 27 x 1980, leg. & det. MJW [FSCA, RWG]; Quijos: 1 ♂, 1 ♀, Baeza 4.8–5.0 km S, small clear streams and seeps (-0.4761, -77.8711, 1830m), 20 xi 1997, leg. & det. WFM [FSCA]; 2 ♂♂, same data but leg. & det. JJD [JJD]; 5 ♂♂, 2 ♀♀, same data but leg. & det. KJT [KJT]; 3 ♂♂, 2 ♀♀, same data but leg. & det. TWD [SWD]; 1 ♀, same data but 10 v 2009, [KJT]; 1 ♂, same data but 3 xi 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, Baeza, 1 mile S on Hwy. 45 (-0.4641, -77.8842, 1830m), 3 xi 2001, leg. & det. F. Sibley [FSCA]; 1 ♂, stream, Baeza-Tena Road, near San Isidore Road, 24 vii 1996, leg. & det. KJT [KJT]; PC, Mejía: 1 ♀, Tatatambo, about 5 km E on Hwy. 30, rocky stream (-0.45, -78.73, 2286m), 30 i 1997, leg. & det. WFM [FSCA]; PA, Mera: 1 ♀, seep near Río Anzu, 12.0 km N of Mera (-1.4181, -78.0528, 1190m), 21 ix 2005, leg. & det. KJT [KJT]; 1 ♂, [Mera] Baños-Puyo Road, [about 5 km W], small forested pond (-1.4806, -78.1444, 1097m), 8 xi 2001, leg. & det. F. Sibley [FSCA]; ZC, Yantzaza: 1 ♀, Yantzaza, 10.8 km N [on Hwy. 45], rocky gravel stream (-3.7669, -78.6915, 762m), 6 xi 1997, leg. & det. WFM [FSCA]; Zamora: 1 ♂, stream/waterfall W of Zamora (-3.9583, -79.0289, 1660m), 1 iv 2008, JTJ [JTJ]; 1 ♀, 30 km E of Loja, foot of cascade, 23 ix 1990, leg. OSF, det.? [USNM].

Macrothemis hemichlora ( Burmeister, 1839) . Publ. Recs. ECU: * Calvert (1906), Ris (1913a), Campos (1922), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14). Also ARG CEN, COL, GUF, PER, SUR, TTO, VEN: (P11); plus BRA: (H06).

Addl. Data. EO, Arenillas: [JTJ, KJT]; Piñas: [ACRP, FSCA]; Portovelo: [FSCA]; LR, Buena Fé: [SWD]; MN, Flavio Alfario: [FSCA]; MS, Morona: [FSCA, KJT]; Santiago: [SWD]; NA, Tena: [DRP, FSCA, UTIC, KJT, LSAM, QZAC, SWD, TWD, USNM]; OR, Francisco de Orellana: [KJT]; SU, Shushufindi: [FSCA, KJT]; ZC: Nangaritza: [FSCA]; Yantzaza: [FSCA].

* Macrothemis idalia? Ris, 1919 . Publ. Recs. GUY, VEN: (H06), (P11).

Addl. Data. OR, Loreto: 1 ♀, 12 km NE of Loreto , forest (-0.6217, -77.2600, 365m), 12 x 2009, leg. KJT, det. RWG [FSCA]; SU, Lago Agrio : 2 ♀♀, [Santa Cecillia]. 24 km W of Lago Agrío on road to Baeza, stream, 20 viii 1980, leg. SWD, det. WFM [FSCA].

Note: Rosser Garrison (pers. comm.) compared the ♀ from near Loreto with ♀♀ in his collection and tenta- tively identified it as this species. A male from Sucumbíos Province, near Shushufindi, collected by KJT on 13 May 2009 was sent to R. W. Garrison who replied “... this could be the unknown ♂? of M. idalia Ris – size, body pattern, claws etc. consistent with known females; but until new material, association would be difficult.” Further collecting, especially a mated pair, will be necessary to confirm this record. Heckman (2006) stated that “without a description of the ♂, the status of this species is uncertain.”

* Macrothemis imitans Karsch, 1890 . Publ. Recs. ECU: (H14). Also ARG, BOL, BRA, CEN, GUY, PRY, TTO, URY, VEN: (H06), (P11); plus COL: (P&P).

Addl. Data. NA, Tena: 3 ♂♂, 1 ♀, Puerto Napo 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde (-1.0500, - 77.7481, 450m), 15 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 3 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 13 vii 1996 [KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. WFM [FSCA, QZAC]; 3 ♂♂, Puerto Napo 11.2 km E of Hwy. 45 on J.S.B.S. Road, Río Tiyuyacu (-1.0516, -77.7017, 396m), 13 xi 1997, leg. & det. WFM [FSCA, QZAC, USNM]; OR, Francisco de Orellana: 1 ♂, [Pom- peya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 229m), 17 xi 1997, leg. & det. WFM [FSCA]; PA, Pastaza: 1 ♂, Río Sandalias , 5 km S of Puyo, Hwy. 45 (-1.5003, -77.9522, 1000m), 11 xi 1997, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. WFM [FSCA]; SU, Cascales: 1 ♂, river, 37 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; Lago Agrio : 1 ♂, stream, 26 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD].

Macrothemis inequiunguis Calvert, 1895 . Publ. Recs. ECU: * Calvert (1898, 1906), Ris (1913a), Campos (1922), Tsuda (1991), (H06), (H14). Also CEN, VEN: (H06), (P11); plus COL: (H06).

Addl. Data. SD: 2 ♂♂, (as M. cf. inequiunguis ), Alluriquín 7.3 km W, forest stream at Hotel Tinalandia, 20 vii 1977, leg. & det. DRP [DRP].

Note: The distribution data indicate that M. inequiunguis is confined to the west slope, its range extending northward to Central America and extreme southern United States, whereas M. tessellata (see below), a very similar species considered by some to be synonymous with M. inequiunguis , is confined to the east slope.

* Macrothemis ludia Belle, 1987 . Publ. Recs. ECU: (H14). Also VEN: (H06), (P11); plus SUR: (P16).

Addl. Data. NA, Tena, 1 ♀, J.S.B.S., near Tena, in garden (450m), 12 vi 1994, leg. A. Jaramillo, det. DRP [DRP]; OR, Francisco de Orellana: 1 ♂, T. B.S., Río Tiputini , 55 km SE of Pompeya, 94 km ESE of Coca (-0.6369, -76.1503), date not provided, data provided by WAH & DLW; SU, Shushufindi: 1 ♂, Limoncocha , forest stream, 9 iii 1972, leg. D. Pearson, det. DRP [DRP].

Macrothemis mortoni Ris, 1913 . Publ. Recs. ECU: (H06), (P11), (H14); NA: * Tennessen (1996b). Also BOL PER, VEN: (H06), (P11).

Addl. Data. NA, Archidona: 1 ♂, stream 43.5 km W of Loreto (1100m), 14 vi 1995, leg. & det. KJT [KJT]; Tena: 1 ♂, stream, road to J.S.B.S., 11.2 km E of Puerto Napo, 15 vi 1995, leg. & det. KJT [KJT]; 1 ♂, Puerto Napo, 3.2 km E on J.S.B.S. Road, sloughs along Napo River, 23 vii 1996, leg. & det. SWD [SWD]; MS, Huamboya: 1 ♂, stream 10 km S. of Comunidad Chuwitayo (-1.9500, -77.8583, 800m), 10 xi 1997, leg. & det. TWD [SWD].

Macrothemis musiva Calvert, 1898 . Publ. Recs. ECU: Schmidt (1942), (H06), (P11), (H14); ES: * Ris (1913a), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, PRY, PER, VEN: (H06), (P11).

Addl. Data. MS, Gualaquiza: 3 ♂♂, 3 ♀♀, Gualaquiza (-3.40, -78.58, 800m), iv–v 1935, leg. F. Campos, det. MJW [ANSP, FSCA]; NA, Tena: 4 ♂♂, 1 ♀, Puerto Napo 3.2 km E on J.S.B.S. Road ponds and flowing marsh (-1.0456, -77.7467, 420m), 23 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC, USNM]; 2 ♂♂, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♀, Puerto Napo 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde (-1.0500, -77.7481, 450 m), 22 vii 1996, leg. & det. WFM [FSCA]; 1 ♀, Puerto Napo 27.7 km E of Hwy. 45 on J.S.B.S. Road, Río Chinchipino (-1.0728, -77.5864, 385m), 13 xi 1997 [FSCA]; 2 ♀♀, same data but leg. & det. KJT [KJT]; 1 ♂, Tena, 13 ix 1990, leg. OSF, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, P.N.Y., 2.3 km E of Yasuni Biological Research Sta., small stream (-0.67, -76.37), 9 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P.N.Y., 3.6 km S of Yasuni Road on Waorani Reserve Road (-0.6956, -76.4594, 244m), 10 vi 1995, leg. & det. WFM [QZAC]; 1 ♂, P.N.Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (-0.6719, -76.3992, 244m), 18 vii 1996, leg. & det. WFM [FSCA]; 1 ♂, stream, Yasuni Road , 20.5 km N Sci. Res. Sta., 11 vi 1995, KJT [KJT]; Loreto: 1 ♂, pond & small stream 34 km W of Loreto (835m), 14 vi 1995, leg. & det. SWD [JJD] 4 ♂♂, Pano , 12 ix 1990, leg. OSF, det.? [USNM]; SU, Cascales: 2 ♂♂, stream 58 km W of Lago Agrio, El Paraiso (455m), 8 vi 1995, leg. & det. KJT [KJT]; Lago Agrio: 2 ♂♂, stream, E edge of Lago Agrio , 18 viii 1980, leg. & det. SWD [SWD]; PA, Pastaza: 1.5 km S of Puyo, 19 v 1977, leg. P. Spangler & D. Givens, det.? [USNM].

Macrothemis pseudimitans Calvert, 1898 . Publ. Recs. ECU: Muttkowski (1910), Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); EO: Campos (1927); ES: Campos (1922); IM: Ris (1913a), Campos (1922); GY: * Calvert (1906), Campos (1922). Also BRA, CEN, COL, TTO, VEN: (H06), (P11); plus PER: (P16).

Addl. Data. IM, Ibarra: 3 ♂♂, “Guayupe” [presumably “Guallupe”] (-0.78, -78.30, 1000m), 13 ii 1946, leg. L. Alonzo, det. MJW [FSCA]; 1 ♂, Lita, 45.8 km E, rocky stream (-0.78, -78.32, 855m), 6 ii 1997, leg. & det. WFM [FSCA].

Note: It is possible that some of the previous records for this species are actually for the very similar M. fallax , which was not described until 1998.

Macrothemis tessellata ( Burmeister, 1839) . Publ. Recs. ECU: * Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14). Also ARG, BRA, CEN, COL, PRY: (P11); plus SUR, VEN: (P16).

Addl. Data. NA, Tena: 1 ♂, Río Sinde, 5 km E of Puerto Napo (-1.0500, -77.7481, 450m), 15 i 1995, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 2 ♂♂, P.N.Y., 3.6 km S of Yasuni Road on Waorani Reserve Road (-0.6956, -76.4594, 244m), 10 vi 1995, leg. & det. WFM [FSCA, QZAC]; 1 ♀, P.N.Y., Yasuni Biological Research Sta. along trails and roads within 5 km of Res. Sta. (-0.6719, -76.3992, 244m), 18 vii 1996, leg. & det. WFM [FSCA]; PA, Mera : 1 ♂, Río Anzu 12.0 km N of Mera, 21 ix 2005, leg. & det. JJD [JJD]; SU, Shushufindi: 1 ♂, trails, small streams 3 km W of Shushufindi, 2+ km N off Hwy. 26 at km marker 17 (-0.1664, -76.6917, 290m), 14 x 2009, leg. & det. JTJ [JTJ]; 1 ♂, Limoncocha , 23 viii 1980, leg. & det. SWD [SWD]; 1 ♂, Shushufindi , about 3 km W, small stream, (-0.1662, -76.6879), 28 x 2013, leg. & det. KJT [FSCA].

Note: We are treating these specimens from the east slope, previously determined to be M. inequiunguis , as M. tessellata ; see also note under M. inequiunguis .

Macrothemis spp. unresolved: IM, Cotacachi: 1 ♀, Santa Rosa , El Refugio de Intag Cloud Forest, south along road at roadside seeps (0.3693, -78.4728, 1854m), 23 viii 2018, leg. & det. WFM [FSCA]; same data but 1 ♂, roadside seeps and open small open stream (0.3703, -78.4745, 1866m), 23 viii 2018 [FSCA]; 2♂♂, 2♀♀, same data but leg. KJT [KJT]; PC, Quito : 1 ♂, 1 ♀, Mindo (1354m), 15 v 2011, leg. & det. ACRP [ACRP]; SU, Shushufindi : 1 ♀, Shushufindi road, 2.5 km W of Shushufindi at km marker 18, 3.5 km S, (-0.2136, -76.6786, 280 m), 12 v 2009, leg. KJT (specimen labeled M. newtoni ?); 1 ♂, 1 ♀, Shushufindi , road, about 3 km NW of Shushufindi, (-0.1664, -76.6167, 290m), 13 v 2009, leg. KJT (both of these specimens were determined by RWG as M. cf. brevidens ).

Miathyria Kirby, 1889

A genus of just two species, easily distinguished ( Ris 1913b). Both species are found in lentic waters such as ponds and canals with floating aquatic plants, especially water hyacinth and water lettuce; the nymphs hide in the roots. See Fig. 101 .

Miathyria marcella (Selys in Sagra, 1857) . Publ. Recs. ECU: Maria (1938), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); CR: Campos (1922); CB: Campos (1922); EO: Campos (1927); GY: * Therese (1900), Calvert (1906), Ris (1913b), Campos (1922, 1930, 1932); LR: Calvert (1906), Campos (1922). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PRY, PER, TTO, URY, VEN: (H06), (P11); plus SUR: (P16).

Addl. Data. BO, San Miguel: [USNM]; CN, Cañar: [FSCA]; EO, Pasaje: [QZAC]; GY, Duran: [ANSP]; Guayaquil: [ANSP, FSCA, USNM]; Naranjal: [FSCA]; LR, Babahoyo: [USNM]; Buena Fé: [DRP]; Ventanas: [JJD]; MN: Chone: [USNM]; Sucre: [USNM]; SU, Shushufindi: [FSCA].

Miathyria simplex ( Rambur, 1842) . Publ. Recs. ECU: Ris (1913b), Paulson (1983), Tsuda (1986, 1991), (H06), (P11), (H14); EO: Campos (1927); GY: * Calvert (1906), Campos (1922, 1930). Also BRA, CEN, GUF, GUY, PER, VEN: (H06), (P11); plus TTO: (P11); and COL, SUR (P16).

Addl. Data. GY, Guayaquil: 1 ♀, Guayaquil, 18 viii 1917, leg. F. Goding, det.? [USNM]; OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza], [across Río Napo from], near Primavera “Limoncocha”, Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 26 vii 1980, leg. & det. SWD [SWD]; 3 ♂♂, P.N.Y., Río Savaleto 2 & slough at km 31, 17 vii 1996, leg. & det. SWD [SWD]; 2 ♂♂, trib. of Río Payamino , road to Loreto, 1.7 km W of jct. with Coca Road (-0.4367, -77.0017, 258m), 18 xi 1997, leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. WFM [FSCA, QZAC]; 1 ♀, Puerto Francisco de Orellana, 11.8 km W on Loreto Road, large pond (-0.4742, -77.0583, 285m), 18 xi 1997, leg. & det. WFM [FSCA]; 1 ♀, same data but leg. & det. KJT [KJT]; SU, Lago Agrio: 1 ♀, 2 km N of Lago Agrio, 27 viii 1975, leg. A. Langley, det.? [USNM]; Shushufindi: 6 ♂♂, 2 ♀♀, Limoncocha , Río Napo (-0.4000, -76.6167, 269m), 6 xi 1980, leg. & det. MJW [FSCA]; 2 ♀♀, same data but 25 xi 1980 [FSCA]; 1 ♀, same data but 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, Limoncocha , clearing, 24 vii 1977, leg. & det. D. Paulson [DRP]; 2 ♂♂, Limoncocha , lakeshore, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 25 viii 1980, leg. & det. SWD [SWD].

Figures

FIGURE 99. Libellula herculea ♂, ECUADOR, Orellana Province, ditch-side stream S of Cascada de Pasahurco, 16.6 km W of Loreto, 4 xi 2013; photo K. J. Tennessen.

FIGURE 100. Macrothemis hahneli ♂, ECUADOR, Napo Prov., small stream <1 km E of Río Sardinas Grande, 24 x 2013; photo K. J. Tennessen.

FIGURE 101. Miathyria marcella ♀, UNITED STATES, Alabama, Fort Morgan Historic Site, 25 xi 2017; photo K. Chias- son.