Micrathyria Kirby, 1889

Mauffray, William F. & Tennessen, Kenneth J., 2019, A Catalogue and Historical Study of the Odonata of Ecuador, Zootaxa 4628 (1), pp. 1-265 : 177-182

Micrathyria Kirby, 1889

A large group of small to medium-sized dragonflies ( Fig. 102 ) in need of revision. Heckman (2006) devised a key to species based on the scattered literature. Twenty-one species have been recorded for Ecuador, plus there are a number of specimens with questionable identity that could increase this number. The group occupies a wide variety of habitats including vegetated lakes, ponds, and sloughs, both open and shaded.

Micrathyria aequalis ( Hagen, 1861) . Publ. Recs. ECU: Muttkowski (1910 —as M. septima and proposed synonymy with M. aequalis ), Paulson (1983), Tsuda (1986, 1991), Silsby (2001), (H06), (P11), (H14); GY: * Calvert (1906), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H06), (P11); plus TTO: (P11).

Addl. Data. EO, Arenillas: 1 ♂, farm pond, seeps, 11 km S of Arenillas, on road to Alamar (-3.6436, - 80.0989, 95m), 10 iv 2008, leg. & det. JTJ [JTJ]; 2 ♂♂, same data but leg. & det. KJT [KJT]; Machala: 1 ♀, borrow pit pond, Hwy. 25, S of Machala (-3.4750, -79.9778), 10 iv 2008, leg. & det. KJT [KJT]; Santa Rosa: 1 ♂, pond, 12 km E of Arenillas, Hwy. 25 (-3.5244, -79.9831, 30m), 29 iii 2008, leg. & det. KJT [KJT]; 3 ♂♂, same data but leg. & det. JTJ [JTJ]; 2 ♂♂, same data but 9 iv 2008, leg. & det. KJT [KJT]; Piñas: 1 ♀, Piedras (-3.63, -79.92, 300m), 6 vii 1941, leg. D. Laddey, det. KJT [FSCA]; GY, Daule: 1 ♂, Nobol, 5.5 km N, (30m), 12 i 1978, leg. P. Spangler & J. Anderson, det.? [USNM]; Guayaquil: 1 ♂, Guayaquil, leg. F. Campos, det. MJW [ANSP]; LR, Babahoyo: adults, 20 km N of Babahoyo, 22 vi 1975, leg. J. Cohen, A. Langley & P. Monnig, det.? [USNM]; Ventanas: 1 ♂, pond/stream 7.2 km E of Hwy. E25, 9 km SW of Quinzaloma (-1.2539, -79.3703, 100m), 21 ii 2010, leg. & det. JTJ [JTJ]; MN, El Carmen: 1 ♂, El Carmen, i 1983, leg. & det.? [QZAC]; 1 ♂, borrow pits about 18 km NW of El Carmen (-0.1350, -79.5797, 185m), 2 iii 2010, leg. & det. JTJ [JTJ].

* Micrathyria artemis Ris, 1911 . Publ. Recs. ARG, BRA, GUF, GUY, PER, SUR, VEN: (H06), (P16).

Addl. Data. NA, Tena: 1 ♂, Puerto Napo, 3.2 km E on Jatun Sacha Road, 23 vii 1996, leg. & det. SWD [FSCA]; SU, Shushufindi: 1 ♂, Limoncocha, lake edge (-0.4000, -76.6167, 269m), 8 ii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP, checked by RWG [DRP].

Micrathyria atra ( Martin,1897) . Publ. Recs. ECU: (H06), (P11), (H14). Also ARG, BRA CEN, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. MS, Santiago: 1 ♂, [Patuca], [about 1 mi N], 43.4 km S of Sucua, large pond, [about 5 km N of Mendez road jct] (-2.75, -78.27, 488m), 7 xi 1997, leg. & det. KJT [KJT]; 1 ♀, same data but leg. & det. WFM [FSCA]; 3 ♂♂, same data but 8 xi 1997 [FSCA, LSAM, QZAC]; 7 ♂♂, 2 ♀♀, same data but leg. & det. JJD [JJD]; 2 ♂♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 1 ♂, forest pool 14.8 km E of Puerto Napo on J.S.B.S. Road, 15 vi 1995, leg. & det. SWD [JJD]; 1 ♂, same data but leg. & det. KJT [KJT]; 1 ♂, Río Sinde, 5.3 km E of Puerto Napo (-1.0500, -77.7481, 450m), 15 vi 1995, leg. & det. KJT [KJT]; 1 cop. pair, Puerto Napo, 14.8 km E on J.S.B.S. Road, forest pool, 15 vi 1995, leg. & det. SWD [SWD]; 2 ♂♂, 1 ♀, J.S.B.S., Pond #14, 18–23 xi 1991, leg. E. McCallie, det.? [USNM]; OR, Loreto: 2 ♂♂, S of Loreto, forest pond, (-0.8141, - 77.3371, 350m), 2 xi 2013, leg. & det. CDK [FSCA]; 1 ♂, N of Loreto, pond, (-0.8205, -77.3202), 2 xi 2013, leg. & det. KJT [FSCA]; SU, Cascales: 1 ♂, pool, 58 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 26 ♂♂, 2 ♀♀, 1 cop. pair, Limoncocha , Río Napo (-0.4000, -76.6167, 269m), 2–26 xi 1980, leg. & det. MJW [FSCA]; 3 ♂♂, 2 ♀♀, same data but palm swamp, 25 vii 1977, leg. & det. DRP [DRP].

Micrathyria caerulistyla Donnelly, 1992 . Publ. Recs. ECU: (H06), (P11), (H14). Also CEN: (H06), (P11).

Addl. Data. MS, Santiago: pond along road between Sucua and Mendez (-2.6833, -78.2283, 600m), 6 xi 1997, leg. & det. TWD [SWD]; NA, Tena: 6 ♂♂, Puerto Napo, 3.2 km E on J.S.B.S. Road, ponds and flowing marsh (-1.0456, -77.7467, 420m), 23 vii 1996, leg. & det. WFM [FSCA, JJD, LSAM, QZAC, USNM]; 4 ♂♂, same data but leg. & det. KJT [KJT]; 4 ♂♂, same data but leg. & det. SWD [RWG, SWD]; 3 ♂♂, 5.3 km E of Hwy. 45 on J.S.B.S. Road, Río Sinde (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♂, same data but 9 x 2009, leg. & det. JTJ [JTJ]; OR, Francisco de Orellana: 5 ♂♂, Puerto Francisco de Orellana, 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. WFM [FSCA, QZAC, USNM]; 1 ♂, 1 cop. pair, same data but leg. & det. KJT [KJT]; 2 ♂♂, same data but leg. & det. SWD [SWD]; 1 ♂, same data but 18 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, same data but leg. & det. TWD [SWD]; 2 ♂♂, [FSCA, JJD]; 1 ♂, 12.3 km W of Coca Road on Loreto Road (820), 13 vi 1995, leg. & det. SWD [SWD]; 1 ♂, P.N.Y., 6.2 km W of Yasuni Road on PC 29-30, pool/stream, 10 vi 1995, leg. & det. SWD [SWD]; 1 ♀, [Pompeya Sur] S on [Pompeya Sur-Iro road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 229m), 17 vii 1996, leg. & det. SWD [SWD]; 1 ♂, P.N.Y., 6.3 km S of Yasuni Road on Waorani Reserve Road, marshy pond (-0.7120, -76.4745, 259m), 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, P.N.Y., 2.1 km E of Yasuni Biological Research Station pond and swamp (-0.6741, -76.3685, 244m), 15 xi 1997, leg. & det. WFM [FSCA]; SU, Shushufindi, 1 ♂, Limoncocha, lakeshore & nearby areas (-0.4000, - 76.6167, 269m), 4 ix 1978, leg. KWK, det. MJW [FSCA]; La Selva Lodge, Lake Mandicocha & outlet stream (-0.4983, -76.3733, 200m), 25 xi 1997, leg. & det. TWD [SWD]; ZC, Yantzaza: 1 ♂, slow stream 7 km N of El Pangui, 6 xi 1997, leg. & det. KJT [KJT].

Micrathyria catenata Calvert, 1909 . Publ. Recs. ECU: (H06), (P11), (H14); ES: Daigle et al. (1997); NA: Tennessen (1996b); SU: * Westfall (1992). Also ARG, BRA, CEN, GUF, PER, SUR, TTO, VEN: (H06), (P11); plus COL: Palacino et al. (2017).

Addl. Data. ES: San Lorenzo: [FSCA, JJD, KJT]; MS, Santiago: [FSCA, JJD, KJT, LSAM, QZAC, TWD, USNM]; Taisha: [DRP]; NA, Archidona: [FSCA]; Carlos Julio Arosemena Tola: [FSCA]; Chaco: (KWN), Tena: [DRP, FSCA, JJD, KJT, SWD, USNM]; OR, Francisco de Orellana: [FSCA, KWK, KJT, LSAM, QZAC, RWG, SWD, TWD, USNM]; SU, Cascales: [SWD]; Shushufindi: [FSCA, KJT]; PA, Pastaza: [FSCA, KJT, QZAC, USNM].

Micrathyria dictynna Ris, 1911 . Publ. Recs. ECU: (H06), (P11), (H14); MS: Tennessen (2006); OR: von Ellenrieder & Garrison (2011); SU * Donnelly (1998). Also CEN, VEN: (H06), (P11); plus COL: (P&P).

Addl. Data. MS, Huamboya: 2 ♂♂, small stream w/ pools, forested, near Río Pasanac [57 km N of Macas], Hwy. 45, N of river, E side of Hwy. (-1.9550, -77.8567), 16 ix 2005, leg. & det. JJD [JJD]; 1 ♂, 1 ♀, same data but leg. & det. KJT [KJT]; NA, Tena: 2 ♂♂, J.S.B.S., pond #13, 23 xi 1991, leg. E. McCallie, det.? [USNM]; 1 ♂, Río Sinde, pond, 5.3 km E of Puerto Napo (-1.0517, -77.7544, 450m), 13 vii 1996, leg. & det. KJT [KJT]; 1 ♂, Río Chinchipino, oxbow pool, 27 km E of Puerto Napo (-1.0728, -77.5864, 385m), 13 xi 1997, leg. & det. KJT [KJT]; OR, Francisco de Orellana: 1 ♀, Yuturi , 2 iii 1990, leg. & det.? [QZAC]; 2 ♂♂, T. B.S., USFQ, Guaca- mayo trail between markers 450–2100 (-0.6350, -76.1533, 237m), 2009, leg. & det. RWG & NVE [NVE, RWG]; Loreto: 1 ♂, forest about 12 km NE of Loreto (-0.6217, -77.2600, 365m), 15 x 2009, leg. KJT [KJT]; same data but 1 ♂, 3 xi 2013, leg. & det. KJT [FSCA]; 1 ♂, S of Loreto, forest pond, (-0.8141 -77.3371, 350m), 2 xi 2013, leg. & det. CDK [FSCA]; PA, Mera: 1 ♂, seepage stream, Los Copales Cabanas between Mera and Shell, W of Puyo, Hwy. 50, 22 ix 2005, leg. & det. JJD [JJD]; SU, Lago Agrio: 1 ♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 1 ♀, Limoncocha, Río Napo (-0.4000, -76.6167, 269m), 11 ix 1973, leg. B. Drummond III, det.? [FSCA]; same data but forest edge, 25 vii 1977, leg. & det. DRP [DRP]; 1 ♂, stream about 3 km NW of Shushufindi (-0.1664, -76.6917, 290m), 13 v 2009, leg. & det. KJT [KJT]; 1 ♂, La Selva Lodge; forest (-0.4983, -76.3733, 200m), 22 xi 1997, leg. & det. TWD [SWD]; 1 ♀, La Selva, Manicocha, 12 x 1988, leg. & det. SWD [SWD]; 1 ♂, 1 ♀, La Selva, Garzacocha, 12 x 1988, leg. & det. SWD [SWD].

Micrathyria didyma (Selys in Sagra, 1857) . Publ. Recs. ECU: Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b as M. didyma didyma ), Silsby (2001), (H06), (P11), (H14); EO: Campos (1927); GY: * Calvert (1906), Ris (1911a), Campos (1922); IM: Ris (1911a—as M. didyma didyma ), Campos (1922). Also CEN, COL, GUF, GUY, SUR, VEN: (H06), (P11); plus BRA: (H06); plus TTO: (P11).

Addl. Data. GY, Guayaquil: 1 ♂, Guayaquil (-2.20, -79.90, 8m), date?, leg. F. Campos, det. P. Calvert, checked WFM [FSCA].

Micrathyria hippolyte Ris, 1911 . Publ. Recs. ECU: (H06), (P11), (H14); NA: * Tennessen (1996b). Also BRA, GUF, GUY, PER, SUR, VEN: (H06), (P11).

Addl. Data. NA, Tena: 1 ♂, [Puerto Napo], [upstream near jct with Río Mulatos] Jatun Yacu Río Napo watershed (-1.05, -77.98, 700m), 25 ii 1937, leg. WCM, det.? [FSCA]; 2 ♂♂, pond, road to J.S. B.S., 14.8 km E of Puerto Napo, 15 vi 1995, leg. & det. KJT [KJT]; 2 ♂♂, 1 ♀, same data but leg. & det. SWD [SWD]; SU, Shushufindi: 1 ♂, Limoncocha (-0.4000, -76.6176, 269m), 28 viii 1980, leg. & det. SWD [SWD]; 1 ♀, same data but forest trail, 27 ix 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♀, same data but 24 i 1972 [DRP]; 1 ♂, same data but 19 ii 1972 [FSCA]; 1 ♂, same data but forest stream, 9 iii 1972 [DRP]; 1 ♂, same data but forest edge, 24 vii 1977, leg. & det. DRP [DRP].

Micrathyria iheringi Santos, 1946 . Publ. Recs. ECU: * Daigle et al. (1996), (H06), (P11), (H14). Also BRA: (H06), (P11).

Addl. Data. MS, Huamboya: 1 ♂, Macas, 58 km N on Hwy. 45 (70 km N of Sucua), roadside ditch and tribs. near Río Pasanac (-1.9447, -77.8536, 762m), 10 xi 1997, leg. & det. WFM [FSCA]; 1 ♂, (labeled “PA”), small stream about 1 km S of Río Pasanac (-1.9503, -77.8631, 820m), 16 ix 2005, leg. & det. KJT [FSCA]; 1 ♂, same data but 16 ix 2005 [KJT]; NA, Tena : 1 ♂, Tena (-0.98, -77.8, 500m), date?, leg. F. Campos, det. KJT [FSCA]; 1 ♂, J.S. B.S., 18 x 1988, leg. & det. SWD [SWD]; 1 ♂, same data but Pond #8, 25 xi 1991, leg. E. McCallie, det.? [USNM]; 2 ♂♂, Río Sinde , slough; 5.3 km E Puerto Napo (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [RWG, SWD]; 1 ♂, same data but 13 vii 1996, leg. & det. SWD [SWD]; 4 ♂♂, pool near Río Napo, 3.2 km E of Puerto Napo (-1.0456, -77.7467, 420m), 23 vii 1996, leg. & det. KJT [KJT]; 1 ♂, same data but leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 17.2 km E on J.S.B.S. Road, pond near stream (340m), 22 vii 1996, leg. & det. SWD [SWD]; 1 ♂, Puerto Napo, 2 nd stream E of Río Sinde on J.S.B.S. Road, 23 vii 1996, leg. & det. SWD [SWD]; OR, Francisco de Orellana: 1 ♂, Comunidad El Rocio, E on Rt E20, pond, (-0.5056, -77.1566), 27 x 2013, leg. & det. KJT [FSCA]; Loreto: 3 ♂♂, Pano , 12 ix 1990, leg. OSF, det.? [USNM]; SU, Shushufindi : 1 ♂, trails, small streams 3 km W of Shushufindi, 2+ km N off Hwy . 26 at km marker 17 (-0.1664, -76.6917, 290m), 13 x 2009, leg. & det. JTJ [JTJ]; ZC, Zamora : 3 ♂♂, Zamora (-4.07, -78.95, 1000m), 21 x 1941, leg. D. Laddey, det. JTJ [FSCA].

Micrathyria laevigata Calvert, 1909 . Publ. Recs. ECU: Ris (1911—as M. didyma laevigata ), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b—as M. didyma laevigata ), (H06), (P11), (H14); SD: Daigle et al. (1997). Also CEN, COL, PER, VEN, TTO, BRA: (H06), (P11).

Addl. Data. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha” Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 4 xi 1980, leg. & det. MJW [FSCA]; La Joya de los Sachas: 2 ♂♂, 2 ♀♀, Pompeya. 5 km W of Puerto Pompeya Ferry Landing, roadside ditches (-0.4444, - 76.6263, 238m), 19 vii 1996, leg. & det. WFM [FSCA, LSAM, QZAC]; 1 ♂, same data but leg. & det. JJD [JJD]; 3 ♂♂, same data but leg. & det. KJT [KJT]; SU, Shushufindi: 3 ♂♂, Limoncocha, Río Napo (-0.4000, -76.6167, 269m), 5 xi 1980, leg. David Robinson & MJW, det. MJW [FSCA]; 2 ♂♂, 4 ♀♀, same data but clearing, forest edge, swamp & lakeshore, 24–26 vii 1977, leg. & det. DRP [DRP].

Micrathyria mengeri Ris, 1919 . Publ. Recs. ECU: * Dunkle (1995), (H06), (P11), (H14). Also BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (P11).

Addl. Data. SU, Shushufindi : 2 ♂♂, Limoncocha, swamp, (-0.4000, -76.6167, 269m), 26 vii 1977, leg. & det. DRP [DRP]; 3 ♂♂, same data but swamp channel, 25 viii 1980, leg. & det. SWD [RWG, SWD].

Micrathyria occipita Westfall, 1992 . Publ. Recs. ECU: (H06), (P11), (H14); NA: Tennessen (1996b); OR: * Westfall (1992, Holotype & Allotype, 1 cop. pair, ”Lake Taracoa near Primavera” = [Francisco de Orellana], [La Nueva Esperanza] [across Río Napo from] near Primavera “Limoncocha” Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 26 viii 1980, leg. SWD [FSCA], Bridges (1993), von Ellenrieder & Garrison (2011). Also PER: (H06), (P11); plus VEN: (P11).

Addl. Data. NA, Tena: [FSCA, UTIC, KJT, QZAC, TWD, USNM]; OR: Francisco de Orellana [DRP, FSCA, JJD, KJT, KWN, LSAM, QZAC, RWG, TWD, USNM]; SU, Lago Agrío: [DRP, FSCA, KJT, USNM]; Shushufindi: [FSCA, KJT, QZAC, RWG, USNM]; ZC, Zamora: [JJD].

Note: An east slope species; specific data for most records can be found in Westfall (1992).

Micrathyria ocellata Martin, 1897 . Publ. Recs. ECU: Muttkowski (1910), Paulson (1983), Tsuda (1986, 1991), Steinmann (1997b—as M. ocellata ocellata ), (H06), (P11), (H14); CR: Calvert (1906), GY: * Martin (1897 — TYPE, Guayaquil, date?); Bridges (1993), Calvert (1906), (Ris, 1911a—as M. ocellata ocellata ), EO: Campos (1927); NA: Tennessen (1996b). Also ARG, BOL, BRA, CEN, COL, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. EO: Machala: [KJT]; Quininde: [KWK, FSCA]; Santa Rosa [JTJ, KJT]; GY, Daule: [USNM]; Guayaquil: [FSCA, USNM]; LR: Quevedo: [FSCA]; MN, El Carmen: [JTJ]; Pedernales: [JTJ]; MS, Santiago: [SWD]; NA, Tena: [FSCA, JJD, KJT, LSAM, QZAC, SWD, USNM]; OR, Francisco de Orellana: [KJT, QZAC]; SU. Lago Agrío: [USNM]; Shushufindi: [DRP, FSCA, KJT, KWN, SWD].

Micrathyria pseudeximia Westfall, 1992 . Publ. Recs. ECU: (H06), (P11), (H14); SU: * Westfall (1992 —description). Also BOL, BRA, CEN, COL, GUF, PRY, PER, SUR, VEN, TTO: (H06), (P11); plus ARG: (P11).

Addl. Data. LR, Buena Fé: 7 ♂♂, 1 ♀, 1 cop. pair, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200m), 18 vii 1977, leg. & det. DRP [DRP]; 8 ♂♂, 1 ♀, paratypes, same data but 8–9 x 1988, leg. & det. SWD [SWD]; MN, El Carmen: 2 ♂♂, borrow pits about 18 km NW of El Carmen (-0.1350, -79.5797, 185m), 2 iii 2010, leg. & det. JTJ [JTJ]; NA, Tena: 1 ♂, Puerto Napo 5.3 km E on Jatun Sacha Road Río Sinde swamp and pond approx. ½ km upstream (-1.0517, -77.7544, 450m), 12 xi 1997, leg. & det. WFM [FSCA]; OR, Francisco de Orellana: 1 ♂, [Pompeya Sur] S on [Pompeya Sur-Iro Road] at km marker 31, borrow pit near Río Sabaleto (-0.60, -76.47, 238m), 17 vii 1996, leg. & det. SWD [SWD]; 1 ♂, pond, road to Waorani Reserve , 6.2 km S of turn to Yasuni Sci . Res. Sta., 10 vi 1995, leg. & det. WFM [FSCA]; 1 ♂, swamp, P.N.Y. (-0.6508, - 76.3558, 200m), 16 xi 1997, leg. & det. KJT [KJT]; 1 ♂, 1 ♀, same data but leg. & det. TWD [SWD]; 1 ♂, stream 2.3 km E of Yasuni Sci. Res. Sta., 9 vi 1995, leg. & det. SWD [SWD]; 1 ♂, Puerto Francisco de Orellana 1.7 km W on Loreto Road from Coca-Sachas Road, stream, pools & flowing shallow marsh (-0.4367, -77.0017, 258m), 13 vi 1995, leg. & det. SWD [SWD]; 1 ♂, pond 8.2 km W of Río Payamino, Coca , 18 xi 1997, leg. & det. JJD [JJD]; 1 paratype ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “Limoncocha”, Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 26 viii 1980, leg. & det. SWD [SWD]; Loreto: 1 ♂, S of Loreto, forest pond, (-0.8141, -77.3371, 350m), 2 xi 2013, leg. & det. CDK [FSCA]; PC, Quito: 2 ♂♂, Pacto Parriquia, San Jose de Mashpi, SE on Mashpi road, small marshy pond & adjacent Tilapia ponds (0.1676, -78.9024, 542m), 3 ix 2018, leg. & det. KJT [KJT, USFQ]; SD: 1 paratype ♂, Santo Domingo de Los Colorados , 12 km E on Hwy. 30 near Hotel Tinalandia, pond with water lilies (-0.30, -79.67, 610m), 6 x 1988, leg. & det. SWD [SWD]; 1 ♂, Al- luriquin, 7.3 km W, Hotel Tinalandia, 11 iii 1990, leg. C. Anderson, det. DRP [DRP]; SU, Lago Agrio : 1 paratype ♂, brushy ponds, 24 km W of Lago Agrio, 20 viii 1980, leg. & det. SWD [SWD]; 2 ♂♂, swamp, 17 km S of Lago Agrio, 19 viii 1980, leg. & det. SWD [SWD]; Shushufindi: 2 ♂♂, Limoncocha, swamp, 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 paratype ♂, same data but 25 viii 1980, leg. & det. SWD [SWD]; 1 paratype ♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (-0.4819, -76.3733, 200m), 12 x 1988, leg. & det. SWD [SWD].

Micrathyria romani Sjöstedt, 1918 . Publ. Recs. ECU: (H06), (P11), (H14); SU: * Dunkle (1981), Westfall (1992). Also BRA, PER: (H06), (P11).

Addl. Data. SU, Shushufindi: 3 ♂♂, Limoncocha, Río Napo (-0.4000, -76.6167, 269m), 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 23 ♂♂, 2 ♀♀, same data but 1–25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but forest edge, 26 vii 1977, leg. & det. DRP [DRP]; 43 ♂♂, same data but lake edge, 26 xii 1971, D. Pearson, det. DRP [DRP]; 16 ♂♂, same data but 26 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 11 vii 1977, leg. & det. KWK [SWD]; 1 ♀, same data but 29 viii 1978, det. JJD [FSCA]; 2 ♂♂, same data but 23 viii 1980, leg. & det. SWD [KJT]; 5 ♂♂, [Limoncocha], [about 40 km E], or 100 km E of Coca [on Río Napo], Lago Garzacocha, La Selva Jungle Lodge (-0.4819, -76.3733, 200m), 11 x 1988, leg. & det. SWD [FSCA, SWD]; 1 ♂, same data but in forest, 6 v 1990, leg. & det. E. Smith [SWD].

Micrathyria spinifera Calvert, 1906 . Publ Recs. ECU: * Paulson(1983), (H06), (P11), (H14). Also BOL, BRA, GUF, GUY, PER, SUR, TTO, VEN: (H06), (P11).

Addl. Data. NA, Archidona: 1 ♂, along Archidona-Loreto Road, stream, pasture to forested, (-0.7164, - 77.6640, 1185m), 25 x 2013, leg. & det. CDK [FSCA]. OR, Francisco de Orellana: 1 ♂, [La Nueva Esperanza] [across Río Napo from], near Primavera “Limoncocha”, Río Napo & Lake Taracoa (-0.45, -76.77, 300m), 27 vii 1977, leg. DRP & S. Hills, det. DRP [DRP]; 1 ♂, same data but 4 xi 1980, leg. & det. MJW [FSCA]; La Joya de los Sachas: 1 ♂, road at Río Napo ferry, S of Limoncocha, 17 vii 1996, leg. & det. KJT [KJT]; SU, Shushufindi: 1 ♂, Limoncocha, Río Napo (-0.4000, -76.6167, 236m), 6 ix 1972, leg. T. Emmel, det. JTJ [FSCA]; 1 ♂, same data but forest trail, 10 viii 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 24 vii 1977, leg. & det. DRP [DRP]; 7 ♂♂, 1 ♀, same data but forest edge, 25–26 vii 1977 [DRP]; 1 ♂, same data but lake edge, 2 x 1971, leg. D. Pearson, det. DRP [DRP]; 1 ♂, same data but 29 viii 1978, leg. KWK, det. JJD [KWK]; 2 ♂♂, Limoncocha, small clearing near Playaco River , 24 viii 1980, leg. & det. SWD [SWD].

Micrathyria sympriona Tennessen, 2000 . Publ. Recs. ECU: (H06), (P04), (P11), (H14); ZC: * Tennessen (2000 — Holotype & Allotype: cop. pair: [Zamora], cutgrass marsh 5.5 km SE of Zamora (-4.17, -78.94, 914m), 5 xi 1997, leg. WFM [FSCA]). Also PER: (H06), (P11); plus COL: Palacino et al. (2017).

Addl. Data. MS, Gualaquiza: 2 ♂♂, 1 ♀, Gualaquiza (-3.40, -78.58, 800m), iv–v 1935, leg. F. Campos, det. WFM [ANSP, FSCA]; ZC, Zamora: 2 paratype ♂♂, Río Jamboe slough 4.2 km SE of Zamora, 5 xi 1997, leg.

JJD, det. KJT [JJD]; 1 paratype ♂, 1 paratype ♀, cutgrass marsh 5.5 km SE of Zamora (914m), 5 xi 1997, leg. JJD, det. KJT [JJD]; 6 paratype ♂♂, same data but leg. & det. KJT [KJT]; 7 ♂♂, same data but leg. WFM [FSCA, LSAM, QZAC, USNM]. 1 ♂, small stream 5 km N of Guaguayme Alto (-3.8778, -78.8928, 880m), 4 iv 2008, leg. & det. JTJ [JTJ].

Note: The Gualaquiza specimens were labeled M. didyma .

Micrathyria tibialis Kirby, 1897 . Publ. Recs. ECU: * Paulson (1983), (H06), (P11), (H14). Also ARG, BOL, BRA, CEN, GUY, PRY, PER, SUR, VEN: (H06), (P11); plus COL: (P&P); and GUF: (P16).

Addl. Data. OR, Francisco de Orellana: 14 ♂♂, 2 ♀♀, Anangu [SE across Río Napo from Providencia], Lago Anangu “on Río Napo ” (-0.5213, -76.4361, 300m), 9 v 1975, leg. B. Drummond III, det.? [FSCA]; SU, Shushufindi: 1 ♀, Limoncocha, Río Napo (-0.4000, -76.6167, 269m), 8 vi 1977, leg. S. Steiner & V. Warren Jr, det.? [USNM]; 13 ♂♂, same data but 3–16 vi 1977, leg. P. Spangler & D. Givens, det.? [USNM]; 1 ♂, same data but 25 viii 1980, leg. & det. SWD; 1 ♂, same data but 25 viii 1980 [SWD]; same data but 6–25 xi 1980, leg. & det. MJW [FSCA]; 1 ♂, same data but lake edge, 22 i 1972, leg. D. Pearson, det. DRP [DRP]; 2 ♂♂, same data but 16 ii 1972 [DRP]; 10 ♂♂, same data but 24–26 vii 1977, leg. & det. DRP [DRP]; 1 ♂, same data but 3 ix 1980, leg. & det. KWK [SWD]; 1 ♂, same data but 25 viii 1980, leg. KWK, det. JJD [FSCSA].

Micrathyria venezuelae De Marmels, 1989 . Publ. Recs. ECU: *(H06), (P11). Also VEN: (H06), (P11); plus (ARG): (P16); and COL: Palacino et al. (2017).

Addl. Data. EO, Portovelo: 1 ♂, Portovelo (-3.72, -79.62, 600–650m), 5 viii 1941, leg. D. Laddey, det. JJD [FSCA]; ES, Quininde: 1 ♂, Rosa Zarate, Río Cupa (0.42, -79.42, 90m), 10 i 1972, leg. D. Poveda, det. JJD [FSCA]; GY, Guayaquil: 1 ♂, Guayaquil (-2.20, -79.90, 8m), date?, leg. N. Banks, det. JJD [FSCA]; LJ, Loja: 1 ♂, seep trib. of Río Chamba, Vilcabamba (-4.2636, -79.2172, 1555m), 31 iii 2008, leg. & det. JTJ [JTJ]; LR, Buena Fé: 1 ♂, Santo Domingo, 47 km S, forest slough at Río Palenque Field Station (200m), 18 vii 1977, leg. DRP, det. WFM [FSCA]; 6 ♂♂, 8–9 xi 1988, leg. & det. SWD [FSCA]; Urdaneta: 1 ♂, ditch/wooded area, 2.8 km E of Hwy. E25 from km marker 98 (-1.5217, -79.5000, 35m), 21 ii 2010, leg. & det. KJT [FSCA]; 2 ♀♀, 5 km E of Hwy. E25 (from Formisa), small stream/secondary forest, (-0.7253, -79.4444), 23 ii 2010, leg. & det. KJT (FSCA); MN, Manta : 1 ♂, Santa Rosa, about 60 km SW of Manta near Río Seco (-1.12, -80.88, 40m), 11 v 1946, leg. L. Alonzo, det. JJD [FSCA]; PA, Mera : 1 ♂, small streams/trails, Los Copales, between Mera and Shell (-1.4919, -78.0725, 1070m), 20 ix 2005, leg. & det. KJT [KJT]; 1 ♂, same data but 22 ix 2005, leg. & det. JJD [JJD]; 1 ♂, Río Anzu , 12.0 km N of Mera (-1.4181, -78.0528, 1190m), 21 ix 2005, leg. & det. KJT [KJT]; SD: 3 ♂♂, seeps at Hotel Colorados , Hwy. 30, about 12 km E of Santo Domingo, 30 i 1997, leg. & det. JJD [JJD]; 2 ♂♂, rocky pool, 8 km W of Alluriquin, Hwy. 30, 31 i 1997, leg. & det. JJD [JJD].

Micrathyria unresolved:

a. Micrathyria cf. almeidai Santos, 1945 . Publ. Recs. ECU: *(H14). Also BRA: (H06), (P16). SU, Shushufindi: 1♂, Limoncocha, no addl. data provided, data provided by WAH& DLW). According to W. A. Haber (pers. comm. 2016), “It likely is not almeidai , although it keys to almeidai in Heckman (2006) , but the appendages are different.”

b. Micrathyria cf. eximia: Kirby, 1897 . Publ. Recs. ECU: (H14). Also COL, ARG, BRA: (H06), (P16). NA, Tena: 1 ♂, Puerto Napo, 5.3 km E, Río Sinde (-1.0500, -77.7481, 450m), 16 x 1988, leg. & det. SWD [RWG]. According to Rosser Garrison (pers. comm. 2011), “Probably a new species.”

c. Micrathyria cf. surinamensis: Geijskes, 1963 . Publ Recs: GUF, SUR: (H06, P16) SU: Shushufindi: 1 ♂, La Selva, 100 km E Coca, Garzacocha (-0.4983, -76.3733, 200m), 12 x 1988, leg. & det. SWD [RWG], data provided by RWG.

d. Micrathyria sp.: ZC, Zamora: 1 ♂, shaded roadside ditch/pool, about 10–11 km N of Guaguayme Alto (-3.8356, -78.8939, 875m), 4 iv 2008, leg. & det. JTJ [JTJ].

e. Micrathyria spp. (H14): “Three different species all in the laevigata group which needs to be revised; one of which is a new species according to Paulson and Garrison” (Haber & Wagmer pers. comm. 2016).

Figures

FIGURE 102. Micrathyria ocellata ♂, COLOMBIA, Meta Dept., pasture pond 5.3 km E of Cumaral, 11 x 2014; photo K. J. Tennessen.