Triaenodes tapanti, Holzenthal & Contents, 2004

Holzenthal, Ralph W. & Contents, Trond Andersen Table Of, 2004, The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae), Zootaxa 511 (1), pp. 1-80 : 34-35

Triaenodes tapanti , new species

Fig. 26 , 37

This new species lacks an apicoventral projection on the inferior appendage, but among similar species the strongly upturned, acute apex of tergum X distinguishes it from all others.

Adult (male: n=10, female: n=4). Forewing length 5.6–6.5, 5.9 mm (male), 6.1–6.6, 6.3 mm (female); hind wing length 4.3–4.9, 4.6 mm (male), 4.8–4.9, 4.9 mm (female). Eye 0.27–0.34, 0.30 mm wide (male), 0.32–0.34, 0.33 mm wide (female). Antennal scape 0.31–0.39, 0.36 mm long (male), 0.34–0.40, 0.37 mm long (female). Maxillary palp segment lengths (in mm): 0.19–0.23, 0.22; 0.31–0.35, 0.33; 0.35–0.39, 0.37; 0.21–0.23, 0.21; 0.44–0.56, 0.47 (male); 0.21–0.23, 0.22; 0.35–0.39, 0.37; 0.44–0.47, 0.45; 0.19–0.24, 0.22; 0.45–0.50, 0.48 (female). Forewing dark golden brown.

Male genitalia. Abdominal segment IX anterior margin broadly rounded, produced anteroventrally; pleural region very lightly sclerotized, heavily setose; tergum IX, in dorsal view, with posteromesal margin rounded, dorsomesal papillae not evident; sternum IX in lateral view with posterior margin rounded. Preanal appendage setose, shorter than wide, broad basally, apex rounded. Dorsomesal process of tergum X digitate, originating basally, long, about half length of tergum X, with apical setae; tergum X, in dorsal view, with lateral margins sinuate, narrowing apically; in lateral view, narrowly triangular, apex acute, strongly upturned, apicolateral margins with microtrichia. Inferior appendage rounded basally, setose, without apicoventral projection; recurved process gradually downturned, narrow, tapering throughout length, but broadest medially, apex attenuate; apicomesal lobe heavily setose, elongate, narrow, parallel sided. Phallobase small, triangular, endothecal membranes prominent, highly convoluted, with three parameres, two parameres subequal, long, straight, tapering, third straight, very slender; phalicata troughlike, strongly curved, narrow throughout length, apex rounded, irregularly serrate, phallotremal sclerite not evident.

Female genitalia. Abdominal segment VIII with sternum setose posteriorly, pleural region highly membranous. Posterior margin tergum IX broadly triangular in dorsal view, with pair of small papillose lobes. Valves heavily setose, rounded, expanded dorsolaterally. Tergum X setose, triangular in lateral view, ventrolateral margin shallowly excavate, apex acute; in dorsal view uniformly wide, apex truncate. Appendage of segment X setose, broad basally, rounded. Structures of internal genitalia as in Figs. 37A, C ; vaginal chamber in ventral view cordate.

Holotype male: COSTA RICA: Cartago: Reserva Tapantí , unnamed tribs [Quebrada Palmitos & falls], ca. 9 km (road) NW tunnel, 09°43’12”N, 083°46’48”W, 1400 m, 8–9.vi.1988, C & O Flint & Holzenthal ( UMSP000083697 ) ( UMSP).

Paratypes: COSTA RICA: Alajuela: Quebrada Virgencita , 10.2 km S Bajos del Toro, 10°10’05”N, 084°19’34”W, 1780 m, 5–6.ix.1990, Holzenthal, Blahnik & Huisman — 1 male ( UMSP) ; Cartago: Reserva Tapantí, Río Grande de Orosí , 09°41’10”N, 083°45’22”W, 1650 m, 15–16.vii.1987, Holzenthal, Morse & Clausen — 1 female ( UMSP) ; Reserva Tapantí , unnamed tribs. [Quebrada Palmitos & falls], ca. 9 km (road) NW tunnel, 09°43’12”N, 083°46’48”W, 1400 m, — 7 males ( UMSP) ; 2–3.vi.1990, Holzenthal, Blahnik & Muñoz — 1 male ( UMSP) ; 1–2.viii.1990, Holzenthal, Blahnik & Muñoz — 1 female ( UMSP) ; 23.viii.1990, Holzenthal & Huisman — 1 female, 1 male ( UMSP) ; Reserva Tapantí , waterfall, ca. 1 km (road) NW tunnel, 09°41’24”N, 083°45’ 36”W, 1600 m, 2–3.viii.1990, Holzenthal, Blahnik & Muñoz — 1 female ( UMSP) ; San José: Zurquí , viii.1995, P Hanson — 1 male ( UMSP) . PANAMA: Chiriqui: Guadalupe Arriba , 08°52’26”N, 082°33’13”W, 7–13.iii.1984, H Wolda — 1 male ( NMNH) ; 11– 17.iii.1984, H Wolda — 1 male ( NMNH) ; 20–26.iii.1985, H Wolda — 1 male ( NMNH) ; 12–18.vi.1985, H Wolda — 1 male ( NMNH) ; 19–25.vi.1985, H Wolda — 1 male ( NMNH) .

Distribution. Costa Rica, Panama.

Etymology. Named for the Reserva Tapantí, Costa Rica, home to this and many other new and interesting species of Trichoptera .

Figures

FIGURE 26. Triaenodes tapanti, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, lateral.

FIGURE 37. Triaenodes tapanti, new species. Female genitalia: A—lateral; B—dorsal; C—ventral.