Triaenodes clauseni, Holzenthal & Contents, 2004

Holzenthal, Ralph W. & Contents, Trond Andersen Table Of, 2004, The caddisfly genus Triaenodes in the Neotropics (Trichoptera: Leptoceridae), Zootaxa 511 (1), pp. 1-80 : 18-19

Triaenodes clauseni , new species

Fig. 9 , 32

Like the former species, this species is similar to T. kilambe , but it differs from both T. chirripo and T. kilambe in the shape of the dorsomesal process and apex of tergum X.

Adult (male: n=7, female: n=2). Forewing length 5.0–5.8, 5.4 mm (male), 5.7–6.2 mm (female); hind wing length 4.2–4.8, 4.5 mm (male), 4.8–5.0 mm (female). Eye 0.35–0.40, 0.38 mm wide (male), 0.37–0.41 mm wide (female). Antennal scape 0.44–0.48, 0.46 mm long (male), 0.41–0.42 mm long (female). Maxillary palp segment lengths (in mm): 0.26– 0.31, 0.28; 0.37–0.42, 0.39; 0.44–0.52, 0.48; 0.24–0.31, 0.27; 0.53–0.58, 0.56 (male); 0.31–0.33, 0.36–0.39, 0.49–0.54, segments IV and V missing (female). Forewing golden brown.

Male genitalia. Abdominal segment IX anterior margin slightly rounded, slightly produced anteroventrally; pleural region very lightly sclerotized, setose; tergum IX, in dorsal view, with posteromesal margin rounded, with pair of small dorsomesal papillae; sternum IX in lateral view with posterior margin slightly sinuate. Preanal appendage setose, at least twice as long as wide, broad basally, apex subacute. Dorsomesal process of tergum X bifid, originating medially, short, less than half length of tergum X, with apical setae; tergum X, in dorsal view, with lateral margins subtriangular, apex strongly excavate medially; in lateral view, parallel sided, apex rounded, slightly excavate subapicodorsally, apicolateral margins with microtrichia. Inferior appendage subquadrangular basally, setose, with short, tapering, apicoventral projection; recurved process gradually downturned, narrow, tapering throughout length, but constricted subapically, apex attenuate; apicomesal lobe heavily setose, elongate, narrow, parallel sided. Phallobase small, triangular, endothecal membranes prominent, highly convoluted, with pair of parameres, parameres subequal, long, gently curved, tapering, apices slightly curved; phalicata troughlike, straight, widest medially, tapering apically, apex rounded, phallotremal sclerite not evident.

Female genitalia. Abdominal segment VIII with sternum setose posteriorly, pleural region highly membranous. Posterior margin tergum IX triangular in dorsal view, with pair of small papillose lobes. Valves heavily setose, elliptical. Tergum X setose, subquadrate in lateral view, ventrolateral margin rounded, apex acute; in dorsal view constricted medially, apex rounded. Appendage of segment X setose, narrow basally, elliptical. Structures of internal genitalia as in Figs. 32A ; vaginal chamber in ventral view circular.

Holotype male: COSTA RICA: Alajuela: Cerro Campana, Río Bochinche trib. 6 km (air) NW Dos Rios, 10°56’42”N, 085°24’47”E, 600 m, 22–23.vii.1987, Holzenthal, Morse & Clausen ( UMSP000083714 ) ( UMSP).

Paratypes: COSTA RICA: Alajuela: Cerro Campana, Río Bochinche trib. 6 km (air) NW Dos Rios , 10°56’42”N, 085°24’47”E, 600 m, 22–23.vii.1987, Holzenthal, Morse & Clausen — 3 males ( UMSP) ; Reserva Forestal San Ramón, Río San Lorencito & tribs., 10°12’58”N, 084°36’25”W, 980 m, 1–4.x.1986, I & A Chacón — 1 male ( UMSP) ; 30.iii.– 1.iv.1987, Holzenthal, Hamilton & Heyn — 2 males ( UMSP) ; Guanacaste: Parque Nacional Guanacaste , ca. 0.7 km N Est. Maritza, 10°57’36”N, 085°30’00”E, 550 m, 31.viii.1990, Huisman & Quesada — 1 male ( INBIO) . NICARAGUA: Jinotega: Cerro Maz , 14°33’00”N, 085°07’00”W, 220 m, 7–10.ix.1997, Maes & Hernández — 1 male ( NMNH) ; Zelaya: Cerro Saslaya , 13°44’00”N, 085°01’00”W, 700 m, iii.1996, Maes & Hernandez — 3 males ( NMNH) . PANAMA: Bocas del Toro: Miramar , 09°00’00”N, 082°15’00”W, 26.xii.1979 – 1.i.1980, H Wolda — 1 male ( NMNH) ; Chiriqui: Fortuna Dam Site nr. Hornitos , 08°55’00”N, 082°16’00”W, 1050 m, 3.viii.–6.ix.1977, H Wolda — 3 males ( NMNH) ; 9–15.xi.1977, H Wolda — 1 male ( NMNH) ; 4.i.–7.iii.1978, H Wolda — 1 female, 1 male ( NMNH) ; 16.viii.–12.ix.1978, H Wolda — 2 males ( NMNH) ; 20– 26.xii.1978, H Wolda — 1 female, 2 males ( NMNH) ; 21–27.iii.1979, H Wolda — 1 male ( NMNH) ; 13.vi.–10.vii.1979, H Wolda — 2 males ( NMNH) .

Distribution. Costa Rica, Nicaragua, Panama.

Etymology. The species is named in honor of Dr. Philip J. Clausen, Curator, University of Minnesota Insect Collection, in recognition of his near life­long commitment to the care and well­being of the collection.

Figures

FIGURE 9. Triaenodes clauseni, new species. Male genitalia: A—lateral; B—dorsal; C—ventral; D—phallus, lateral.

FIGURE 32. Triaenodes clauseni, new species. Female genitalia: A—lateral.