Jalapyphantes, AND

Moreira, Thiago Da Silva & Hormiga, Gustavo, 2021, Systematics of the Neotropical spider genera Jalapyphantes and Selenyphantes and the circumscription of the Pocobletus clade (Araneae: Linyphiidae), Zoological Journal of the Linnean Society 192 : -

JALAPYPHANTES AND SELENYPHANTES REVISION 933

Spermathecae falciform, with variable orientation and ( Fig. 37). Fertilization ducts short, exiting caudally ( Figs 23G, 25G, 27G, 29G).

Male palp patella with a long, strong macroseta (absent in S. longispinosus ). Palpal tibia ventrally bent, with two to three long, strong macrosetae ( Figs 22H, 24H, 25H, 28H, 30H, 31H); two mesal and two dorsal trichobothria. Cymbium mesally oriented, round and broader on its posterior part than its anterior part (basket shape). Cymbium with an ectobasal projection oriented upwards (above the paracymbium in ectal view) and a mesobasal projecting downwards under the cymbium. Cymbial lobe over the subtegulumbasal haematodocha area. Paracymbium ‘J’-shaped (cymbium contact area is longer than the distal area, with an acute inward angle; Figs 22D, 24D, 26D, 28D, 29D, 30D), not parallel to the tegular area, but projecting outwards (somewhat in angle, or almost orthogonal in frontal view; Fig. 31F) the palp. Paracymbium bears basal and apical setae. Tegular area is larger than the cymbium. Tegular retrolateral lobe (TRL) present, large with variable morphology. Tegulum with perforation area of variable size, within a fold of the tegulum between the TRL and the apex on dorsal view ( Fig. 40C–E). Tegulum apex, projecting outwards and with variable morphology and discrete protegular process, with papillae. Tegulum bears a hook-shaped structure (of difficult observation) close to its ectal margin ( Figs 41F, G, 42C). Suprategulum orthogonal to the tegulum ( Fig. 40G). Suprategular apophysis small, of variable morphology, with no apophysis. SA parallel to the tegular apex ( Figs 22E, 24E, 26E, 28E, 29E, 30E), never surpassing. SA separated from the suprategulum by a weakly sclerotized and transparent area ( Figs 22G, 24G, 26G, 28G, 29G, 30G). Embolic division completely mesal oriented concerning the overall palp. Column basal on suprategulum, thick and exposed ( Figs 22H, 24H, 26H, 28H, 29H, 30H). Radix small, with embolic connection partially membranous. Embolus long (approx. 1.5– 2.0× the length of the prosoma), thick at the basis and gradually thinning through its extension. Embolus

938 T. SILVA-MOREIRA and G. HORMIGA

exits radix into a downward curve ( Figs 34A–C, 35B, C) except in S. costaricensis where it exits in an upward curve; Fig. 35A), projecting ectally, winding counterclockwise.

Embolic division with two non-radical sclerites: the lamella characteristica (LC) and the radical complex (RC; see definition under Jalapyphantes genus description; Figs 32A–C, 33A–C, 34A–C, 35A–C). LC large, ‘comma’-shaped and sclerotized ( Figs 24H, 25H, 28H, 29H, 30H). RCm and RCs (except on S. orizabae and in S. iztactepetl where RCs is absent) of approximately the same thickness and same orientation, curved inward, never projecting dorsally (upwards) ( Figs 24F, 26F, 28F, 29F, 30F).

Natural history: There is no information about the ecology or behaviour of Selenyphantes species.

Distribution: COSTA RICA: Cartago, Puntarenas, San José; GUATEMALA: Alta Verapaz; Quetzaltenango; MEXICO: Distrito Federal, Hidalgo, México, Veracruz.