Exocelina anggiensis Shaverdo, Hendrich & Balke

Shaverdo, Helena V., Surbakti, Suriani, Hendrich, Lars & Balke, Michael, 2012, Introduction of the Exocelina ekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 250, pp. 1-76 : 4-5

2. Exocelina anggiensis Shaverdo, Hendrich & Balke sp. n. Figs 9 A–E, 35

Type locality.

Indonesia: West Papua Province: Manokwari Regency, Arfak Mts., Anggi, Iray, 01°20.51'S, 133°55.64'E.

Type material.

Holotype: male "Indonesia: Papua, Anggi, forest above Iray 1, 2000m, 2.ii.1994, 01.20.512S, 133.55.642E, Balke (BH 06)" (ZSM). Paratypes: 5 males, 8 females with same labels as the holotype, one of them additionally with a green label "DNA M.Balke 1272" (NHMW, ZSM). 1 female "Indonesia: Papua, Anggi, Iray, 1880m, 1.ii.1994, 01.18.224S, 133.54.009E, Balke (BH 05)", "DNA M.Balke 1271" [green] (ZSM).

Diagnosis.

Beetle middle-sized, reddish-brown to brown; submatt; pronotum with lateral bead; male antennomeres 3-4 strongly enlarged and triangular (3 larger than 4), 5-6 distinctly enlarged, 7 somewhat enlarged; male protarsomere 4 with small (only slightly larger than more laterally situated large seta), thin, slightly curved anterolateral hook; median lobe with strong submedian constriction in ventral view, apex of median lobe elongate in lateral view and narrower in ventral view; paramere with notch on dorsal side and subdistal part short and small, with not numerous, relatively short, thick, and flattened setae.

Description.

Size and shape: Beetle middle-sized (TL-H 3.9-4.3 mm, TL 4.25-4.7 mm, MW 2.05-2.3 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Head red to reddish-brown, with brown to dark brown areas behind eyes and “V” -like spot in middle, pronotum red to reddish-brown, with two darker areas laterally on disc (left and right from middle line), in some specimens jointed in middle, elytra reddish brown to dark brown, with reddish lines along sutura, head appendages yellowish red to red, legs red to reddis-brown, slightly darker distally; two paratypes with more uniform brown coloration (Fig. 35).

Surface sculpture: Whole dorsal surface with dense and coarse punctation (spaces between punctures 1-3, often 1-2, times size of punctures). Punctation slightly sparser on elytra (spaces between punctures 1-4 times size of punctures). Pronotum and elytra with strongly impressed microreticulation, dorsal surface, thus, submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal sternites with distinct microreticulation, strioles, and fine, rather dense punctation, coarser and denser on two last abdominal sternites.

Structures: Pronotum with distinct lateral bead, sides somewhat more rounded. Base of prosternum and neck of prosternal process with distinct ridge, not smooth and slightly rounded anteriorly, without anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal sternite 7 broadly rounded apically.

Male: Antennomeres 3-4 strongly enlarged and triangular (3 larger than 4), 5-6 distinctly enlarged, 7 somewhat enlarged (Fig. 9A); antennomeres 3-6 rugose ventrally. Protarsomere 4 with small (only slightly larger than more laterally situated large seta), thin, slightly curved anterolateral hook. Protarsomere 5 ventrally with anterior row of 9 short setae and posterior row of 7 short setae (Fig. 9B). Abdominal sternite 7 with 8-10 lateral striae on each side. Median lobe with strong submedian constriction in ventral view, apex of median lobe elongate in lateral view and narrower in ventral view (Figs 9C, D). Paramere with notch on dorsal side and subdistal part short and small, with not numerous, relatively short, thick, and flattened setae (Fig. 9E).

Female: Dorsal punctation denser and coarser than in males, antennae simple, abdominal sternite 7 without striae.

Distribution.

Indonesia: West Papua Province: Manokwari Regency. The species is known only from the type locality (Fig. 50).

Etymology.

The species is named for the type area, Anggi. The name is an adjective in the nominative singular.