Ectospinula Opitz, 2016

Opitz, Weston, 2016, Classification, Natural History, and Evolution of the Korynetinae (Coleoptera: Cleridae). Part III. The African GeneraEctospinulaOpitz, New Genus, andProsymnusLaporte, The Coleopterists Bulletin 70 (3), pp. 429-446 : 429-446

Ectospinula Opitz , new genus

Type Species. Ectospinula graduata Opitz , new species.

Diagnosis. The gradual increase in the width of the distal antennomeres of the funicle ( Fig. 3 ) will distinguish the members of this genus from others in Korynetinae .

Description. Size: Length 8.0 mm; width 3.5 mm. Form: Oblong, hind body suboval, robust, deep body, about 2.5 times longer than broad. Vestiture: Disc of cranium and pronotum vested with long, dark setae, elytral disc vested with

22) P. astrictus .

epipleural fold wide, ventrally positioned and expanded in proximal half, ends just before elytral apex, outer margin of elytral apex crenulated, anterior margin carinate; metathoracic wing as in Fig. 1 , wedge cell closed; tibial spur formula 2-2-2, tarsal pulvillar formula 3-3-3, unguis with well-developed denticle. Abdomen: With 6 visible sternites, sternites 3 and 4 without setal patches.

Distribution. This monotypic genus is known only from Zambia.

Etymology. The generic name Ectospinula is a compound name derived from the Greek ekto (= without) and the Latin spinula (= dim). I refer to the slightly developed spine near the posterior angles of the pronotum. Gender: feminine.

1° setae and shorter, profusely distributed 2° setae. Head: Cranium quadrate, frons very wide, indented with large, setiferous punctation, latter nearly contiguous; gula small, rectangular, sutures vertical, gular process short, narrow, and forked; labrum short, medial incision deep, curvate concave; mandibular body short, stout; maxillary terminal palpomere subsecuriform; labial terminal palpomere digitiform ( Fig. 5 ); eyes moderately sized, coarsely faceted, ocular notch large; antenna capitate ( Fig. 3 ), capitulum lax, scape large, longer than combined length of pedicel and antennomeres 1–3, basal 3 funicular antennomeres oblong, distal 3 funicular antennomeres progressively increasing in diameter, capitular antennomeres not compacted, antennomeres 9 and 10 transverse, antennomere 11 subovoid. Thorax: Pronotum subovoid, convex, lateral margins minutely crenulated, with a small spine near pronotal hind angle, coarsely sculptured with round, setiferous punctation, prebasal fissure deep, prointercoxal process very expanded distally; pronotal projections long, but do not contact prointercoxal process; elytron sculptured with large, spheroid, asetiferous punctation, latter subseriate, 1° setae always adjacent to asetiferous punctation, 2° setae present and somewhat arranged serially,

Ectospinula graduata Opitz , new species ( Figs. 1, 3, 5 , 19 , 36 , 37 )

Holotype. ♀. Mongu, Barotseld., 1952, J. M. Niemeyer ( Zambia, Mongu , Barotseland) ( TMSA).

Diagnosis. The shape of the funicular antennomeres ( Fig. 3 ) will distinguish the members of this species from any other in Korynetinae .

Description. Size: Length 8.0 mm; width 3.5 mm. Form: As in Fig. 19 . Color: Cranium and pronotum black with a shiny blue luster, antennae and legs black, elytra black with a cupreous luster, abdomen testaceous. Head: Antennal capitulum lax ( Fig. 3 ), antennomeres 9 and 10 transverse and cupuliform, 11 subovoid; last labial palpomere digitiform ( Fig. 5 ); eyes much narrower than frons (EW/FW 18/62). Thorax: Pronotum transverse (PW/PL 145/120), lateral margins serrulated; elytral punctation large to subseriate, interstitial spaces smooth and shiny; elytral apex subacuminate. Abdomen: Setal patches on 3 rd and 4 th sternites; female pygidium scutiform.

Distribution. This species is known only from Zambia ( Fig. 37 ).

Etymology. The specific epithet graduata is a Latin adjective that stems from gradus (= step). The name describes the gradual increase in width of the distal funicular antennomeres.

Figures

Figs. 1–8. Body structures. 1) Ectospinula graduata, metathoracic wing. Antennae: 2) Necrobia rufipes; 3) E. graduata; 4) Prosymnus rudis. Terminal labial palpomere: 5) E. graduata; 6) P. rudis. Abdominal sternites: 7) P. rudis; 8) Prosymnus mulleri.

Figs. 19–22. Ectospinula and Prosymnus species, habitus. 19) E. graduata; 20) P. adustus; 21) P. arsus;

Fig. 36. Phylogenetic tree computed from the character matrix (Table 1) for Ectospinula graduata and Prosymnus species.

Figs. 37–40. Geographic distribution of Ectospinula graduata and Prosymnus species.