Mahellus distanti Nielson, 1982

Meshram, Naresh M., 2019, Leafhopper tribe Coelidiini (Hemiptera: Cicadellidae: Coelidiinae) of the Indian subcontinent, Zootaxa 4653 (1), pp. 1-91 : 18

Mahellus distanti Nielson

Figs. 28 A–F.

Mahellus distanti Nielson 1982: 86 , Figs. 264−269; 2015: 35; Viraktamath & Meshram 2017: 273 , 279, Figs. 3−7 , 11−16 , 18−31 , 48−49 , 51 .

Material examined. INDIA: Maharashtra: 1♂, 4♀, Matheran , 915 M, 23.xi.1977, C. A. Viraktamath . Karnataka: 1♂, 22 Km, S. Bangalore , 22.v.1977, C. A. Viraktamath ; 2♂, Hunsur , 18.i.1978, C. A. Viraktamath ; 2♂, Banerghat- ta, 22.v.1977,C. A. Viraktamath; 1♂, 1♀, Chickballapur , 01.x.2015, M. Shobharani ; 1♀, 24 Km, W. Mudigree , 21.ix.1978, C. A. Viraktamath . Tamil Nadu: 1♀, 64 Km , E Kollegal, 9.viii.1977, C. A. Viraktamath ; 1♀, Shevroy Hills , 21.ix.1978, C. A. Viraktamath . Kerala: 1♂, 1♀, Maraiyur , 24.iii.1977, C. A. Viraktamath ( USAB) .

Remarks. Nielson (1982) adequately described and illustrated the species based on males from Nilgiri Hills. Viraktamath and Meshram (2017) provided digital photos of the habitus and male and female genitalia of the species in addition to giving new locality records. M. distanti differs from the remaining three species of the genus in having a more robust aedeagal shaft and the processes adpressed to the shaft ( Figs. 28 D–F) compared to the more slender aedeagal shaft and the processes oriented away from the shaft in the other species ( Figs. 28 K–M).

Figures

FIGURES 3 A–O. Habitus of Coelidiini species.A. Singillatus serratispatulatus sp. nov., female; B & C. Trinoridia dialata sp. nov., male and female. D. T. ochrocephala sp. nov., male; E & F. T. piperica sp. nov., male and female; G & H. T. ramamurthyi sp. nov., male and female; I. T. saraikela sp. nov., male; J. T. timlivana sp. nov., male; K. T. tripectinata (Nielson), male; L & M. Webbolidia andamana sp. nov., male and female; N. W. burmanica sp. nov., male; O. Zhangolidia weicongi sp. nov., male.

FIGURES 4 A–O. Habitus of Coelidiini species, lateral view. A. Calodia deergha sp. nov., male; B. C. keralica sp. nov., male; C & D. C. kodikanelensis Nielson, male and female; E. C. kumari sp. nov., male; F & G. C. neofusca sp. nov., male and female; H & I. C. ostenta (Distant), male and female; J. C. periyari sp. nov., male; K. C. subcrista Nielson, male; L. C. tridenta sp. nov., male; M. Glaberana acuta sp. nov., male; N & O. G. khasiensis (Rao), male and female.

FIGURES 5 A–O. Habitus of Coelidiini species, lateral view. A. Glaberana purva sp. nov., male; B & C. Olidiana brevis (Walker), male and female. D. O. fletcheri sp. nov., male; E. O. indica (Walker), male; F & G. O. kirkaldyi (Nielson), male and female; H. O. lanceolata sp. nov., male; I. O. perculta (Distant), male; J. O. serra (Neilson), male; K & L. O. umroensis sp. nov., male and female; M. O. unidenta sp. nov., male; N. Singillatus parapectitus sp. nov., male; O. S. serratispatulatus sp. nov., male.

FIGURES 6 A–O. Habitus of Coelidiini species, lateral view. A. Singillatus serratispatulatus sp. nov., female; B & C. Trinoridia dialata sp. nov., male and female. D. T. ochrocephala sp. nov., male; E & F. T. piperica sp. nov., male and female; G & H. T. ramamurthyi sp. nov., male and female; I. T. saraikela sp. nov., male; J. T. timlivana sp. nov., male; K. T. tripectinata (Nielson), male; L & M. Webbolidia andamana sp. nov., male and female; N. W. burmanica sp. nov., male; O. Zhangolidia weicongi sp. nov., male.

FIGURES 7 A–P. Faces of Coelidiini species. A. Calodia deergha sp. nov., male; B. C. keralica sp. nov., male; C & D. C. kodikanelensis Nielson, male and female; E. C. kumari sp. nov., male; F & G. C. neofusca sp. nov., male and female; H & I. C. ostenta (Distant), male and female; J. C. periyari sp. nov., male; K. C. subcrista Nielson, male; L. C. tridenta sp. nov., male; M. Glaberana acuta sp. nov., Male; N & O. G. khasiensis (Rao), male and female. P. Glaberana purva sp. nov., male.

FIGURES 11 A–P. Female valvulae I and II. Calodia neofusca sp. nov.: A & B. Valvula I; C & D. Valvula II. C. ostenta (Distant): E& F. Valvula I; G & H. Valvula II. C. kodikanelensis Nielson: I & J. Valvula I; K & L. Valvula II. Glaberana khasiensis (Rao): M & N. Valvula I; O & P. Valvula II.

FIGURES 12 A–P. Female valvula I and II. Olidiana brevis (Walker): A & B. Valvula I; C & D. Valvula II. O. kirkaldyi (Nielson): E& F. Valvula I; G & H. Valvula II. O. umroensis sp. nov.: I & J. Valvula I; K & L. Valvula II. Singillatus serratispatulatus sp. nov.: M & N. Valvula I; O & P. Valvula II.

FIGURES 13 A–P. Female valvula I and II. Trinoridia dialata sp. nov.: A & B. Valvula I; C & D. Valvula II. T. piperica sp. nov.: E & F. Valvula I; G & H. Valvula II. T. ramamurthyi sp. nov.: I & J. Valvula I; K & L. Valvula II. Webbolidia andamana sp. nov.: M & N. Valvula I; O & P. Valvula II.

FIGURES 14 A–N. Species of Calodia, male genitalia. A–D. C. bicompressa Nielson; E–G. C. fusca (Melichar); H–K. C. paraostenta Nielson; L–N. C. propennata Nielson. Modified and reproduced from Nielson 2015 (Figs. A–D), Nielson 1982 (Figs. E–G) and Nielson 1991 (Figs. H–N).

FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 16 A–H. Calodia keralica sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 18 A–G. Calodia kumari sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view.

FIGURES 19 A–H. Calodia neofusca sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 20 A–H. Calodia ostenta (Distant), male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 21 A–G. Calodia periyari sp. nov., male genitalia. A. Aedeagus, dorsal view; B & C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view.

FIGURES 22 A–I. Calodia subcrista Neilson, male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view; I. Pygofer, ventral view.

FIGURES 23 A–H. Calodia tridenta sp. nov., male genitalia.A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 24 A–V. Species of Creberulidia Nielson and Hiatusorus Nielson, male genitalia. A–D. Creberulidia excelsa (Distant); E–H. C. pica (Nielson); I–L. C. quadrispinata (Nielson); M–Q. Hiatusorus concavus (Nielson); R–V. H. hardyi (Nielson). Modified and reproduced from Nielson 1982 (Figs. A–H, M–Q), Nielson 1991 (Figs. I–L) and Nielson 2015 (Figs. R–V).

FIGURES 25 A–H. Glaberana acuta sp. nov., male genitalia. A. Aedeagus, dorsal view; B. Aedeagus, lateral view; C. Subgenital plate, dorsal view; D. Style, dorsal view; E. Connective, dorsal view; F. Pygofer, lateral view; G. Pygofer dorsal view; H. Pygofer dorsal process, dorsal view.

FIGURES 26 A–H. Glaberana khasiensis (Rao) comb. nov., male genitalia. A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer dorsal process, lateral view; H. Pygofer, lateral view.

FIGURES 27 A–H. Glaberana purva sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view; H. Pygofer posterior extension, lateral view.

FIGURES 28A–M. Species of Mahellus Nielson, male genitalia. M. distanti Nielson: A. Pygofer, lateral view; B. Subgenital plate, ventral view; C. Style, dorsal view; D. Aedeagus, lateral view; E.Aedeagus dorsal view; F. Apex of aedeagal shaft, dorsal view. M. ungulatus Viraktamath & Meshram: G. Pygofer, lateral view, H. Pygofer process, lateral view; J. Style dorsal view; K. Aedeagus, lateral view; L. Aedeagus, dorsal view; M. Apex of aedeagal shaft, dorsal view. Modified and reproduced from Viraktamath & Meshram (2017).

FIGURES 29 A–G. Olidiana brevis (Walker), male genitalia. A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view.

FIGURES 30 A–H. Olidiana fletcheri sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 31 A–G. Olidiana indica (Walker), male genitalia. A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Style, dorsal view; F. Connective, dorsal view; G. Pygofer, lateral view.

FIGURES 48 A–G. Trinoridia saraikela sp. nov., male genitalia.A & B. Aedeagus, dorsal view; C. Aedeagus, lateral view; D. Subgenital plate, dorsal view; E. Connective, dorsal view; F. Styles (asymmetrical) and connective, dorsal view; G. Pygofer, lateral view.

FIGURES 49 A–H. Trinoridia timlivana sp. nov., male genitalia. A & B. Aedeagus, dorsal view; C & D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.

FIGURES 51 A–H. Webbolidia andamana sp. nov., male genitalia.A–C. Aedeagus, dorsal view; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view.