Liogenys, Guerin-Meneville, 1831

Cherman, Mariana Alejandra, Basílio, Daniel Silva, Mise, Kleber Makoto, Frisch, Johannes & Almeida, Lúcia Massutti De, 2021, Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species, Zootaxa 4938 (1), pp. 1-59 : 3-4

publication ID

https://doi.org/10.11646/zootaxa.4938.1.1

publication LSID

lsid:zoobank.org:pub:074E95CD-E7C0-4098-93F0-70DE03CAD0D0

DOI

https://doi.org/10.5281/zenodo.4590912

persistent identifier

https://treatment.plazi.org/id/CD475E66-EC07-5A7F-6BBF-194FFA36FAB5

treatment provided by

Plazi

scientific name

Liogenys
status

 

Identification key to Liogenys species from the Chacoan province and its boundaries

1. Antenna with 10 antennomeres; clypeal emargination wide or narrow; outer sides of anterior clypeal teeth parallel ( Figs. 2B View FIGURE 2 , 3B View FIGURE 3 ) or following the lateral margin of clypeus ( Figs. 9B View FIGURE 9 , 21B View FIGURE 21 ).................................................. 2

- Antenna with 9 antennomeres; clypeal emargination narrow and semicircular ( Fig. 12E View FIGURE 12 ); outer sides of anterior clypeal teeth following the lateral margin of clypeus.......................................... Liogenys mendozana Moser, 1918

2. Clypeal teeth very short ( Fig. 4B View FIGURE 4 ), outer sides of teeth not forming an angle with the lateral margin of clypeus; lateral margin convex or straight, not produced......................................................................... 3

- Clypeal teeth long or short ( Figs. 2B View FIGURE 2 , 8B View FIGURE 8 ), outer sides of teeth forming an angle with the lateral margin of clypeus, lateral margin convex and sometimes strongly produced ( Fig. 6B View FIGURE 6 )....................................................... 8

3. Pronotal disc densely punctate ( Fig. 3B View FIGURE 3 )................................................................... 4

- Pronotal disc sparsely punctate ( Figs. 2B View FIGURE 2 , 10C View FIGURE 10 ).............................................................. 6

4. Pronotum and elytra glabrous; punctures of pronotum very fine and dense ( Fig. 4B View FIGURE 4 ); pygidium with dense scales ( Fig. 4D View FIGURE 4 ), scales sparse only at the apex...................................................... Liogenys densata Frey, 1969

- Pronotum and elytra setose; punctures of pronotum coarse and dense ( Fig. 3B View FIGURE 3 ); pygidium with sparse bristles and scales... 5

5. Clypeal emargination semicircular, narrow, lateral margin of clypeus never produced; in males inner margin of metatibia concave subapically ( Fig. 16F View FIGURE 16 )................................................ Liogenys opacicollis Fairmaire, 1892

- Clypeal emargination rounded, wide, lateral margin of clypeus concave, produced in most of the cases, forming a blunt projection ( Fig. 3B View FIGURE 3 ); in males inner margin of metatibia convex subapically ( Fig. 3F View FIGURE 3 ).......... Liogenys cribricollis Moser, 1921

6. Lateral margin of clypeus concave, in males tarsomeres almost equal in width in all legs, cylindrical ( Fig. 9E View FIGURE 9 )............................................................................... Liogenys isotarsis Cherman , new species

- Lateral margin of clypeus straight ( Fig. 21B View FIGURE 21 ), in males protarsomeres and mesotarsomeres enlarged.................... 7

7. Elytra setose, shiny; body length less than 8 mm ............................. Liogenys truncata Cherman , new species

- Elytra glabrous, semiopaque; body length 9–10 mm .................................. Liogenys rectangula Frey, 1969

8. Pronotum roughly punctate; lateral sides of abdomen, propygidium, and pygidium covered by wide scales ( Fig. 20C View FIGURE 20 )................................................................................. Liogenys rugosicollis Frey, 1969

- Pronotum somewhat coarsely punctate; sides of abdomen, propygidium, and pygidium with bristles ( Fig. 10D View FIGURE 10 )........... 9

9. Width of clypeal teeth combined three or more times longer than the length of one tooth ( Figs. 14B View FIGURE 14 , 18B View FIGURE 18 ).............. 10

- Width of clypeal teeth combined less than three times longer than the length of one tooth ( Figs. 6B View FIGURE 6 , 8B View FIGURE 8 )............... 17

10. Epipleuron of elytra uniformly clothed by fringed-like, long, erect bristles ( Fig. 18A, E View FIGURE 18 ); legs with tarsomere II twice as long as tarsomere I, or almost so, especially in male....................................... Liogenys parallela Frey, 1965

- Epipleuron of elytra with sparse, short bristles; legs with tarsomere II less than twice as long as tarsomere I............. 11

11. Pygidium subquadrate ( Fig. 14D View FIGURE 14 ), apex angulate, punctures very coarse; clypeal emargination deeper than the outer margin of anterior teeth ( Fig. 14B View FIGURE 14 )...................................................... Liogenys nigrofusca Moser, 1918

- Pygidium trapezoidal ( Figs. 2 View FIGURE 2 D–E; 5G–H; 15G), apex subangulate or rounded; clypeal emargination shallower than the outer margin of anterior teeth ( Fig. 10C View FIGURE 10 )....................................................................... 12

12. Lateral margin of clypeus weakly produced, outer sides of anterior teeth obtuse or subparallel, without angle among anterior teeth and lateral margin of clypeus ( Fig. 15E View FIGURE 15 ).............................................................. 13

- Lateral margin of clypeus strongly produced, outer sides of anterior teeth almost parallel, right or acute angle between anterior and lateral teeth ( Fig. 10C View FIGURE 10 )............................................................................. 14

13. Propygidium scaly; pygidium with erect and thick setae............................ Liogenys obscura Blanchard, 1851

- Propygidium with sparse bristles; pygidium with decumbent bristles.................... Liogenys cartwrighti Frey, 1969

14. Pygidium glabrous, or with few bristles on apex; distance between teeth of each tarsal claw as long as the length of superior tooth.............................................................................................. 15

- Pygidium setose, distance between teeth of each tarsal claw shorter than the length of inferior tooth................... 16

15. Body shiny, head and pronotum unicolored brown or dark brown in both sexes; clypeal emargination narrow, without clear sexual dimorphism in the clypeal margin; males with protarsomere II longer than wide ( Fig. 5J View FIGURE 5 )............................................................................................... Liogenys denticeps Blanchard, 1851

- Body opaque, head and pronotum dull yellow unicolored in males, not unicolored in females, head dark brown or black; clypeal emargination very wide, in females emargination as wide as one half the distance between eyes, in males wider; males with protarsomere II as long as wide or wider than long ( Fig. 10D View FIGURE 10 )........................ Liogenys pallens Blanchard, 1851

16. Metafemur glabrous, row of bristles along its margins; males with parameres angulate at the midline, without subapical constriction ( Fig. 11 View FIGURE 11 )............................................................. Liogenys latitarsis Moser, 1918

- Metafemur setose throughout; males with parameres globose at the midline, with subapical constriction ( Fig. 22E View FIGURE 22 )....................................................................................... Liogenys vicina Frey, 1969

17. Clypeal emargination angulate ( Fig. 8B View FIGURE 8 ); fovea of distal maxillary palp occupying the entire dorsal surface, almost reaching the apex ( Fig. 8C View FIGURE 8 )......................................................... Liogenys foveata Cherman new species

- Clypeal emargination rounded; fovea of distal maxillary palp extending past the midline but not reaching the apex....... 18

18. Base of pygidium less than twice as wide as the apex ( Fig. 7E View FIGURE 7 ); in males, the three pairs of tarsi equally enlarged...................................................................................... Liogenys tarsalis Moser, 1921

- Base of pygidium twice as wide as the apex ( Figs. 7F View FIGURE 7 , 13D View FIGURE 13 ); in males, protarsi and mesotarsi wider than the metatarsi.... 19

19. Base of metacoxae and trochanter setose ( Fig. 13E View FIGURE 13 ); in males, parameres glabrous ( Fig. 13F View FIGURE 13 ).......................................................................................... Liogenys neoforcipata Cherman , new species

- Base of metacoxae and trochanter almost glabrous ( Fig. 6D View FIGURE 6 ); in males, parameres setose ( Fig. 6F View FIGURE 6 )... Liogenys forcipata Frey

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Melolonthidae

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