Eurygaster testudinaria (Geoffroy, 1785)

Neimorovets, Vladimir, 2020, Review of the genus Eurygaster (Hemiptera: Heteroptera: Scutelleridae) of Russia, Zootaxa 4722 (6), pp. 501-539 : 525-533

Eurygaster testudinaria (Geoffroy, 1785)

Figs. 18 , 25 , 31 , 37 , 44–49 , 55 , 86–111 .

Cimex testudinarius Geoffroy in Fourcroy, 1785: 195 . For Cimex no. 3 in Geoffroy, 1762: 435.

Eurygaster maura f. borealis Péneau, 1911: 174 (syn. China, 1927: 253, as Eurygaster testudinaria (Geoffroy, 1785)) .

Eurygaster sinicus Walker, 1867: 67 (downgraded by Josifov & Kerzhner, 1978: 166, as Eurygaster testudinaria sinica Walker, 1867 ), syn. nov.

Material examined. RUSSIA: Kaliningrad Prov.: 1♂, Zelentsovo , 13.viii.1954, Levin leg. Berezhkovskoye Le- snichestvo, 3♂, 4♀, Zelentsovo , 4, 5, 6, 8, 10, viii.1954, Levin leg. 1♀, Krasnolesye , 13.vii.1954, Levin leg. Leningrad Prov.: 1♀, Near Kingisepp [Yamburg], 3.vi.1908, D. Glazunov leg. 2♂, 1♀, Khimosi near Gatchina, 5, 10.viii.1940, Unknown collector. 1♂, 1♀, Gatchina , 5.viii.1940, Stackelberg leg. 8♂, 11♀, Gorelovo , 13, 23 , 31. vii.1898, 2, 3, 5, 10, 15, 25.viii.1898, Novotortsev leg. 2♀, Gobzhitsa , 12.vii.1934, 16.viii.1934, Stackelberg leg. 1♂, Luga , 28.viii.1898, Novotortsev leg. 1♂, 1♀, Tolmachyovo , Luzhsky District, 8,14 . viii.1935, Rodendorf leg. 1♀, Luzhsky District [former Gdov uezd] , 14.vii.1907, N. Ivanov leg. 1♀, Rakovichi near Luga , 3–13.viii.1897, Pleske leg. 1♂, 1 nymph 3 th instar, 1 nymph 5 th instar, Lomonosovsky, District, Lebyazhye , 15.viii.1897, vii.1900, Bianki leg. 1♀, Seryozino , 10.viii.1895, Bianki leg. 1♂ Bologoe , 2.vii.1903, Unknown collector. 2♂, Near Pavlov- sk, VIR , 15.vii.1966, Unknown collector. Pskov Prov.: 1♀, Bezhanitsy , 11.vi.1915, Unknown collector. 2♂, Pok- rovskoe, 23.vii.1915, 4.viii.1915, N. Kuznetsov leg. Novgorod Prov.: 1♀, Oxochi , 30.vi.1926, I. Gudim leg. Arkhangelsk ProV.: 2♂, Near Kotlas , 14 , 29.viii.1942, Stark leg. Yaroslavl Prov.: 1♂, 1♀, Danilovsky District, 1915, A. Shestakov leg. 1♀, Zhedenovo 12 km South to Danilov, Danilovsky District, A. Shestakov leg. Kostroma Prov.: 1♀, Mamturovsky District, Ugri , 19.ix.1983, E.M. Veselova leg. Bryansk Prov.: 3♂, 3♀, Bryansk, 14, 17 , 23.vii.1927, Bryansk, 1♂, Kiritshenko leg. 1♀, Bryansk District, “ Bryanskoe opitnoe lestnichestvo”, Vinogradov- Nikitin leg. 4♀, Bryansk , 7.vi.1912, 3.viii.1925, 22.vii.1926, 23.vii.1927, V. Stark leg. Kaluga Prov.: 3♂, 1♀, Ta- rusa, 18 , 19.viii.1956, 21.vi.1961, Lyubishchev leg. Moscow Prov.: 2♂, 1♀, Klinsky District, Boblovo , 4, 7 , 10. vi.1905, Unknown collector. 1♀, Leninsky District, Sukhanovo , 15.vii.1906, A.P. Datskevich leg. 1♀, Stupinsky District, Mikhnevo , 6.viii.1905, B.K. Hindze leg. 1♂, Stupinsky District, Mikhnevo , 6.viii.1905, G.A. Kozhevnikov leg. 1♀, Dmitrovsky District, Ivanovskye [former Dmitrovsky Uezd, Ivanovo] , 24.viii.1908, Shaposhnikov leg. 1♀, Dmitrovsky District, Semenkovo , 15.viii.1908, Shaposhnikov leg. 1♂, 1♀, Mozhaisk District, Porechye , 15.vii.1903, Bianki leg. 1♀, Ruzsky District , 29.vii.1901, N. V. Voronkov leg. 1♂, 1♀, Podolsk, 1912, Letichev leg. Ryazan Prov.: 2♂, Shatsky District, Kiritshenko leg. 1♂, Shatsky District , 1907, Malishev leg. 1♂, Gremyachka , Milo- slavsky District [former Dankovskiy Uezd] , 27.ii.1901, A. Semenov leg. Nizhny Novgorod Prov.: 8♂, 5♀, Staraya Pustin , 13, 17, 18, 22, 23, 24, 25, 27 , 30.viii.1939, Kiritshenko leg. Ulyanovsk Prov.: 1♀, Ulyanovsk, Academgoro- dok , 22.viii.1963, A. Lubischew leg. 1♀, Ulyanovsk , 28.ix.1964, A. Lubischew leg. 1♀, Ulyanovsk, “ Bely Kluch ” , 19.vii.1954, A. Lubischew leg. 1♀, Ulyanovsk , 13.vi.1952, A. Lubischew leg. 2♀, Vinnovskaya Roscha [former Vinnovka], Ulyanovsk , 15.vi.1956, 1♂, 6.viii.1951, A. Lubischew leg. 1♂, 2♀, Ulyanovsk, floodplain of Sviyaga River , 21.vi.1956, A. Lubischew leg. 1♀, Ulyanovsk, Verkhny Zotovsky Island , 6.ix.1952, A. Lubischew leg. 1♂, Ulyanovsk, Seredish Island , 12.viii.1951, A. Lubischew leg. 1♀, Ulyanovsk, Popov Island , 4.vi.1954, A. Lubischew leg. 1♀, Ulyanovsk, Palitsinsk Island , 14.viii.1951, A. Lubischew leg. 1♂, Beloye Lake , 23.viii.1963, 2♀, A. Lubischew leg. 4♀, Ulyanovsk, Zasviyazhsky District , 19.viii.1956, 18.viii.1961, A. Lubischew leg. 6♂, 7♀, Kirzyat, near Glubokoye Lake , 27.vii.1957, 16, 25.viii.1957, A. Lubischew leg. 1♂, 1♀, Surskoye , 18 , 26.vi.1953, A. Lubischew leg. 2♂, “ Zakharyevsky mine” , 25.viii.1952, A. Lubischew leg. 1♂, 2♀, Barysh , 22, 28 , 29.vi.1955, A. Lubischew leg. 1♂, 3♀, Staraya Maina , 19.v.1954, 11, 12.vi.1954, A. Lubischew leg. Tatarstan Rep.: 1♂, Ka- zan, 1907, M. Sizov leg. Kursk Prov.: 1♂, Lgovsky District, Zakharkovo , 12.ix.1898, E. Mertens leg. Belgorod Prov.: 1♀, Borisovka , vii.1990, I. Smetanin leg. 1♀, Borisovka , 18.vi.1907, S. Malishev leg. 1♂, Shebekino [for- mer Shebek, Kurskaya Gubernia] , 29.vi.1900, Serebryannikov leg. 1♂, Grayvoron , 5.ix.1904, S. Malishev leg. Voronezh Prov.: 5♀, Voronezhskoye lesnichestvo, floodplain of Khoper River , “Tellermanova Rosha” , 16.vi.1952, 12, 17.vi.1953, A. Moravskaya leg. 1♀, Gribanovsky District [former Polyanskiy District], Polyanskoye Lesnich- estvo , 29.vii.1953, Stark leg. 1♂, Kamennaya Step, 11 km South to Talovaya , 12.vi.1949, Stark leg. 10♂, 9♀, Ter- novsky District, Svalskoye lesnichestvo , 25.v.1951, 25.v.1953, 1.vii.1953, 4.viii.1953, 11.v.1954, 6, 21.vi.1954, 17.vi.1955 19.vii.1955, Stark leg. 1♂, Ternovsky District, Svalskoye lesnichestvo , 2.vi.1955, Speranskaya leg. 3♂, 2♀, Voronezh , 31.vii.1917, 1♀. Kalatch , 5 , 17.vi.1933, 23.vi.1934, 23.vii.1934, I. Gudim leg. Tambov Prov.: 1♂, Michurinsk [former Kozlov], Unknown date, N. Lange leg. 1♂, Tambov Prov. , 23.viii.1909, B. Grigoryev leg. Penza Prov.: 2♂, Penza, Unknown date, Olsufiev leg. Saratov Prov.: 1♂, 2♀, Kamyshin , 30.v.1920, 20.viii.1920, Mordvilko leg. 1♂, Volsk , 1.viii.1928, V. Vinogradova leg. 2♂, Volsk, 1923, P.S. Kozlov leg. 2♂, 3♀, Khvalynsk , 25.vii.1952, 3, 4.viii.1952, A. Lubischew leg. Samara Prov.: 2♂, 1♀, Samara, Krasnoglinsky City District, Kras- naya Glinka , 30.vi.1929, 15, 24.viii.1929, Lyubishchev leg. 1♀, Volga River , near Samara , 16.viii.1930, Lyubishchev leg. 1♀, Syzran , 28.v.1906, V. Bianki leg. 1♀, Sergiyevsky District, Pavlovka , 21.vii.1907, Elagir leg. 1♂, Samara Prov. , 9.vi.1905, Steklova leg. 1♀, Kinel , 8.ix.1928, Lyubishchev leg. 2♂, 1♀, Zhiguli Nature Reserve , 23.vi.1940, E. Novoderezhkin leg. 1♀, Zhiguli , 23.viii.1928, Dmitriev leg. 1♀, Zhiguli , 24.vi.1926, E. Novoderezhkin leg. Orenburg Prov.: 1♂, Ashchebutak 60 km South to Orenburg , 8.viii.1933, L. Zimin leg. 1♀, Buzuluksky Bor Na- tional Park, Orenburgsky District [former Chkalovsky District] , 18.viii.1941, Chistovsky leg. Bashkortostan Rep.: 1♂, 1♀, Zilim River , 27–28.vi.1925, Gekker leg. 1♂, Burzyansky District, Irgizli [former Irgizla] , 11.vi.1899, Ja- cobs and Schmidt leg. Volgograd Prov.: 1♂, Volgograd, Sarepta [now district of Volgograd], 1872, Bekker leg. 1♂, Volgograd, Sarepta [now district of Volgograd], 1872, Jakovlev leg. 1♀, Kamyshin , 8.vi.1936, Berezina leg. 1♀, Kamyshin District , 17.vii.1950, A. Moravskaya leg. Rostov Prov.: 1♀, Novocherkassk , 25 Mar, Kizertsk leg. 1♂, 2♀, Persianovka , 3.vi.1915, Unknown collector. Astrakhan Prov.: 1♂, Vishka 80 km to South Astrakhan , 16.vii.1961, Kerzhner leg. 1♂, 1♀, Oranjerei , 14.v.1911, Lukash leg. Crimea Rep.: 2♂, Near Alushta , “Korbekli” , 3.viii.1900, N. Kuznetsov leg. 2♂, Near Simferopol [former Post Station Tauman–Bazar] , 7.vi.1907, Grigoriev leg. 2♂, 1♀, Near Simferopol [former Post Station Tauman–Bazar] , 10.vii.1907, Pliginskiy leg. 1♀, Near Simferopol, Dzhalman, Unknown date, Kakhovsky leg. Krasnodar Ter.: 1♀, Unknown locality [NW Caucasus], Stark leg. 1♀, Unknown locality [“Kuban Oblast”], Vorobiev leg. Mostovskoy Distr., 1♂, Caucasus Nature Reserve , kordon “Tre- tya Rota” , 6.vi.2002, V. Neimorovets leg. 4♂, 1♀, near Psebay , 14.vi.2002, V. Neimorovets leg. 1♀, Krasny Les, 1910, N. Bryansky leg. 3♀, Near Ilsky , 13 , 15.ix.2001, V. Neimorovets leg. 1♀, Seversky District, Derbentskaya , 31.v.1998, V. Neimorovets leg. 26♂, 21♀, Near Novodmitrievskaya [the place former named “Shebskaya Voisko- vaya Dacha”], 14 , 26.v.1907, 28.vi.1907, 14, 17, 26, 28.vi.1910, 1, 2, 6.vii.1910, Anderson & Gurov leg. 3♀, Nebug , 16.viii.1925, Gitterman leg. 1♂, 1♀, Uch–Dere near Lazarevskoye, E. König leg. 1♂, 3♀, Lazarevskoye , 31.vii.1952, 2, 9, 19.viii.1952, 2L5, Rudolf leg. 1♀, Thagapsh ( Bozhi void) , 14.vii.1910, I. Smirnov leg. 1♂, Sochi , 10–12.vii.1900, Bikov leg. 1♂, Adler , 27.vii.1907, Kiritshenko leg. 3♂, 6♀, Krasnaya Polyana , v.1907, 8, 10, 11, 15, 18, 21, 26.vii.1907, Kiritshenko leg. Adygea Rep.: 1♀, Maykop , 16.vii.1927, G. Semenov leg. 1♀, Bambak Mount , 17–18.vi.1907, N. Vorobyev leg. 1♀, Upper course of Laba River , 26.vi.1912, Shaposhnikov leg. Stavropol Ter.: 2♀, Mashuk Mount , 10 , 28.vi.1927, Ryabov leg. Karachay– Cherkess Rep.: 2♂, 2♀, Dombay–Ulgen Mount , 2 , 9.viii.1909, Bryansky leg. Kabardino– Balkar Rep.: 1♂, Teberda , 11.viii.1953, L. Arens leg. 1♂, 1♀, Nalchik , 14.iv.1949, Bogachev leg. 1♂, 6♀, Nalchik , 4, 5, 7, 8 , 12.vii.1914, E. Golovlieva leg. 1♂, 2♀, Nalchik, Goluboye Lake , 13.viii.1917, Bartenev leg. North Ossetia – Alania Rep.: 1♂, Vladikavkaz , 6.vi.1920, Unknown collector. 1♂, 1♀, Vladikavkaz , 11.vii.1925, Kiritshenko leg. 2♂, Vadikavkaz , 4 , 30.vii.1909, Vinogradov and Nikitin leg. Ingushetia Rep.: 2♂, Assa River near Alkun , 2.vi.1886, Ananov leg. 1♂, River Martan ( Martanka ) , 1.vi.1886, S.K. Ananov leg. 1♀, Aki–yurt [former Akiyurtovskaya] , 28.v.1886, Ananov leg. Dagestan Rep.: 1♂, Botlikh , 28.vi.1913, A. Mlokosevich leg. Kurgan Prov.: 2♀, Orlovka , 18.viii.1946, Dubinin leg. Tyumen Prov.: 1♂, 4♀, near Gor- noslinkino, Irtysh River , 12.vi.2004, A.B. Rivkin leg. Tomsk Prov.: 1♀, Semiluzhki , 23.v.1908, A. Emelyanov leg. 1♀, Tom River, Near Tomsk , 15.vi.1901, Shafir leg. Novosibirsk Prov.: 1♀, Novosibirsk , 12.ix.1926, Levitsk leg. Kemerovo Prov.: 1♂, 3♀, Teba, Kuznetsk Alatau , 11.viii.1962, Miski, 26.v.1956, 2, 10.vi.1956, Falkovich leg. 1♂, Kamzas Riva, tributary of Tom River , 11.vi.1908, Khvorov leg. 1♂, 5♀, Vaganovo , 31.v.1955, 18.vi.1955, 24.vii.1955, 10, 22.viii.1955, Falkovich leg. Altai Ter.: 1♂, 1♀, Barnaul ( Lebyazhye City District ) , 6.viii.1948, Nikolskaya leg. 2♂, 1♀, Slavgorodsky District [former Slavgorodsky Uezd ] , 22–23.vii.1922, A. Reichard leg. 9♂, 8♀, Barnaul, Altai Research Institute of Agriculture , 15.viii.2014, Unknown collector ( VIZR). 3♂, Barnaul, Altai Research Institute of Agriculture , spring soft wheat , 10.viii, 20.viii.2015, G.G. Sadovnikov leg. ( VIZR). 2♂, 1♀, Tigereksy Nature Reserve , 23.vi.2005, A. Namyatova leg. 3♀, Tigereksy Nature Reserve , 7, 8 , 10.vii.2009, A. Knyshov leg. 5♂, 2♀, Pavlovsk , 28.v.1944, 7.vi.1944, Stark leg. Altai Rep.: 1♂, 3♀, Onguday , 5.viii.1904, 8.viii.1905, V. Steinfeld leg. 2♂, 2♀, Turochak , 19, 24 , 28.vi.1926, D. Suvorov leg. 2♀, North coast of Lake Telets- koye , 11.vi.1901, Ignatov leg. Khakassia Rep.: 1♀, Kuznetskiy Alatau Range, Grunin leg. 2♂, Balyksu River, tributary of Tom River , 9 , 11.vi.1908, Khvorov leg. 2♂, 3♀, Askiz, Abakan River , 5.vii.1925, Unknown collector. Krasnoyarsk Ter.: 1♂, Konny Island on Yenisey River near Krasnoyarsk , 10.ix.1905, Kibort leg. 1 nymph 5 th instar, Near Krasnoyarsk , 14.vi–1.vii.1902, Salstrem leg. 1♂, 1♀, Tukhtat [former Tukhtatka] , 29.v.1901, Kibort leg. Bazarikha, Degtyaryov leg. 1♂, Immiskoye, Kazyr River , 1.vii.1936, Lukyanovic leg. 1♀, Nizhnye Kuryati, Kazyr River , 2.vii.1936, Lukyanovic leg. 1♀, Station Kozulka 66 km East to Achinsk , 22.vii.1936, Lukyanovic leg. 1♂, 1♀, Minusinsky District, Yenisey River , 9.ix.1930, Lukyanovic leg. 1♂, Taseyevo , 17–30.vii.1915, Borisov leg. 6♂, 4♀, Motorskoye , 18, 19 , 20.vi.1925, Serebryanikov leg. 1♂, 1♀, Lesosibirsk [former Maklakovo], 15.vii–1.viii.1915 , 10.ix.1915, Malikh leg. Tyva Rep.: 1♀, Near Chadan, Chadan River , 2.vii.1947, Fedorova leg. 1♀, Uyuk River, tributary of Bolshoy Yenisei River [former Bii–Khem] , 9.vii.1949, Cherepanov leg. 1♂, Turan, Uyuk River , 9.viii.1949, Perevozchikova leg. 1♂, Turan , 5.vi.1949, 1949, 1♀, Perevozchikova leg. 3♀, Turan , 29.viii.1948, Cherepanov leg. 2♀, Chaa–hol River , 2.viii.1948, Cherepanov leg. 1♀, Saryg–Sep, Little Yenisei River [former Ka–Khem] , 15.viii.1948, Kalugin leg. Irkutsk Prov.: 1♀, near Irkutsk , 21.vi.1924, Vinogradov leg. Irkutsk, 1.viii.1912, Mokievskaya leg. 42♂, 47♀, Bunbui , 4–8, 14.vi.1915, 16, 18, 23–30.vii.1915, 2, 11, 13, 17, 18.viii.1915, Varaksina leg. 75♂, 92♀, 7 nymph 5 th instar Bunbui , 6 , 23.vi.1915, 16.vii.1915, 2, 3, 11, 13, 17, 18.viii.1915, 8, 30.vi.1916, 30.vii.1916, Valdaev leg. 2♂, 5♀, Irkutsk , 1898 V. Jakovlev leg. 1♂, Kirensk , 28.v.1904, 1♂, Braudo leg. Yurty, 31.vii.1912, Mishin and Verkhov leg. Buryatia Rep.: 1♀, Kyakhta [former Troitskosavsk] , 23.vi.1928, Luky- anovic leg. Sakha Rep.: 2♂, Ulahan–Botuobuya River [former Bolshaya Batubia River], tributary of Vilyuy River , 27–29.vi.1916, I. Bogomolov leg. 1♂, Between Yakutsk and Amga [former Amga–Yakutsk Tract] , 13.viii.1925, A. Ivanov leg. 1♀, Between Yakutsk and Vilyui, Badaranakh , 16.viii.1926, Ivanov leg. Zabaykalsky Ter.: 5♂, 6♀, 1♂, 1♀, 1♂, Sretensk , 23, 24 , 27.vi.1928, 8, 13.ix.1928, 29.viii.1930, Kapustin leg. 1♂, Ust–Urluk, Selenga River , 4. vii.1928, Lukyanovic leg. 1♀, Bayan [fomer Boyan–Khosun ] , 18.viii.1928, Lukyanovic leg. 1♀, Zharcha River , “Spassky mine” , 1–5.viii.1912, Shmelin leg. 2♀, Ingoda River , viii.1898, A. Kiritshenko leg. 1♂, Darasun [former Darasun–Tiret] , 18.viii.1925, Vinogradov leg. Amur Prov.: 1♂, Ignatyevo , 16.viii, Unknown collector. 2♀, Gomelev- ka, 14.vi., Zavitin leg. 1♂, 2♀, Nikolayevsk–on–Amur [former Nikolayevsk], 1920 , 17.ix.1926, 10.viii.1928, Ros- tovykh leg. 2♂, 1♀, Ignatyevo [former Ignatyevka] , 4.ix.1928, Formozov leg. 2♂, 1♀, the Tambovka River , 26.vi.1915, Soldatov leg. 1♂, Zea River , 50 km near Blagoveshchensk , 8.vi.1914, Popov leg. 1♀, Andreevka , 4.vii.1907, V. Ivanov and V. Soldatov leg. 1♀, Pokrovka, 1908, L.A. Jakovlev leg. 1♂, 4♀, Ulanginskoye , 10, 15 , 16.vi.1910, 30.vii.1910, 4.viii.1910, Mishin and Verkhovsky leg. 1♂, Between Khingan and Ussuri, vi–vii.1957, Radde leg. 1♀, Romanovka , 1.vii.1905, V. V. Barovsky leg. 6♂, 5♀, Salokachi River (left tributary of Arkhara River ), 8, 10 , 18.ix.1946, L. Kiritshenko leg. 1♂, 1♀, 1♀, Between Malaya Pera and Bolshoi Ergel , 11.vi.1957, 14, 27. vii.1957, 1, 9, 18, 19.viii.1957, Zinovyev leg. 1♀, Klimoutsy 40 km West to Svobodny , 29.v.1957, Zinovyev leg. 9♂, 8♀, Klimoutsy 40 km West to Svobodny , 17, 18, 18 , 30.vi.1959, Kerzhner leg. 30♂, 13♀, Simonovo 75 km West to Svobodny , 5, 8, 9, 12, 17, 19 , 23.vi.1959, 11, 13.viii.1959, Kerzhner leg. 3♂, 3♀, Selemdzhinsky District, Norsk, floodplain of Selemdzha River, Nora Nature Reserve , kordon Dvadtsatikha , 18 , 23.vi.2005, Veselova and Rivkin leg. 1♀, Selemdzhinsky District, Norsk , 7.iv.2004, Rivkin leg. 2♀, Tolbuzino, upper course of Amur River , vii.1929, 1– 19.vii.1929, Prinada leg. 1♀, Valley of Ivanovka , 15.viii.1926, Gekker leg. 7♂, 5♀, Novostepanovka , 22.viii.1959, Grunin leg. Khabarovsk Ter.: 1♀, Khabarovsk , 23.vii.1916, Pavlenko leg. 1♂, near Khabarovsk , 25–26.v.1913, V. K. Soldatov leg. 2♂, 3♀, Hechzir Range , ix.1930, Prinada leg. 1♂, Hechzir Range , 26.viii.1979, Ermolenko leg. 1♂, 7♀, Nizhnetambovskoe , 7, 9 , 29.vi.1911, 30.vii.1911, V. Soldatov leg. 4♂, 7♀, near Nizhnetambovskoe , 11 , 12.vi.1911, V. Soldatov leg. 1♀, Kulku River near Nizhnetambovskoe , 30.vii.1911, V. Soldatov leg. 1♂, 1♀, near Khabarovsk, Ussuri River, 1910, V. Soldatov leg. 1♀, Vyatskoye , [former Vyatsk] , 12.vi.1910, V. Soldatov leg. 2♂, 2♀, Near Litvintsevo , 17 , 25.vi.1911, V. Soldatov leg. 3♂, 1♀, Verkhnebureinsky District, Talandga, Burea River , 18, 19, 25 , 27.viii.1946, L. Kiritshenko leg. 1♂, 1♀, 10 km to Bolen, Sumniur River , 21 , 22.vi.1911, A. Chersky leg. 1♂, 2♀, near Korsakovo , 25.vii.1959, 4, 7.viii.1959, Kerzhner leg. 1♂, 1♀, 30 km NW to Pobeda, Kur River , 30.ix.1978, Kabakov leg. 1♂, Nizhnetambovskoe , 20.viii.1926, Starokodom leg. 1♂, 1♀, Kazakevichevo , 11.vii.1925, Unknown collector. Jewish Autonomous Prov.: 1♂, Kuldur River (tributary of Bira River ) , 31.viii.1946, L. Kiritshenko leg. 1♂, Bira , 16–17.vi.1931, Prinada leg. 3♂, 4♀, Birobidzhan, Tikhonkaya Station , 20–22.vi.1931, 30.vi.1931, 2.vii.1931, Prinada leg. Primorsky Ter.: 1♂, Primorsky Territory [former Ussuri Treeitory], Jakovlev leg. 1♀, Vlad- ivostok , 20.viii.1909, Derbek leg. 1♂, 2♀, Vladivostok , 29.vi.1911, 5, 27.vii.1911, Ridzovsky leg. Vladivostok, viii.1926, N.N. Pegoaotski leg. 1♀, Vladivostok , 18.vi.1909, Berger and Chersky leg. 1♂, 1♀, Vladivostok, Sedanka , 8.ix.1909, Berger leg. 1♀, near Vladivostok , 1.ix.1910, Barisheva leg. 1♀, near Vladivostok, Okeanskaya Station , 10.vii.1959, Falkovich leg. Anisimovka [former Kangauz ] , 15.vii.1927, A. Lorvi leg. 1♂, 1♀, middle course of Arte- movka River [former Mayhe] , 12.vii.1930, Shambovsky leg. 1♂, near Shkotovo , 5.vi.1927, Sokolov leg. 1♀, near Shkotovo , 5.vi.1927, Stackelberg leg. 1♂, Tigrovoy [former Tigrovoe] , 16.vii.1927, Stackelberg leg. 1♂, 1♀, Yakov- levka, 1927, Kvashuk leg. 2♀, Yakovlevka , 25.v.1927, 27.vi.1927, Stackelberg leg. 1♀, Mouth of Armu River, tribu- tary of Bolshaya Ussurka River [former Iman River] , 16.viii.1946, Nesmerchuk leg. 1♂, Bolshaya Ussurka River [former Iman River] , 15–28.vii.1931, Shablinovsky leg. 1♂, 3♀, Dalnerechensk [former Iman], 5 , 15.v.1911, 13.vi.1911, 20.vii.1911, Shingar leg. 1♀, Dalnerechensk [former Iman] , 13.vi.1899, V. Jakovlev leg. 2♂, Evseyevka , 2.vii.1910, Ikonnikov leg. 1♀, middle flow of Ussuri River [former Ulahe] , 20.vi.1902, Bunin leg. 1♂, lower flow of Arsenyevka River [former Daubikhe] , 24.v.1911, Shingarev leg. 1♀, Nizhnemihaylovka , 16.vi.1899, Suvorov leg. 3♂, 1♀, Beshenaya River , 4, 8 , 14.vii.1911, V. Soldatov leg. 1♂, 1♀, 1 nymph 5 th instar, Troitskoye , 27–28.vi.1908, 20.vii.1909, Chersky leg. 1♂, Kamen–Rybolov, Khanka Lake , 5.vi.1908, Dukin leg. 1♂, 3♀, Kamen–Rybolov, Khanka Lake , 27.vi.1908, 1.ix.1908, Chersky leg. 3♂, 3♀, Kamen–Rybolov, Khanka Lake , 18.vi.1910, 3, 9.vii.1910, Tarabarov leg. 1♂, near Khanka Lake , 2.vii.1909, A. Chersky leg. 2♂, 2♀, 20 km East to Khanka , 24.vi.1910, Ikon- nikov leg. 12♂, 11♀, Jakovlevka , 10.iv.1926, 17, 19, 25, 28.vi.1926, 30 13.vii.1926, viii.1926, 1, 2, 3, 8, 10, 11, 12.ix.1926, Dyakonov and Filipyev leg. 1♂, Vityaz, 15 km south to Sukhanovka , 8.ix.1982, Kerzhner leg. 1♂, 2♀, Bezverhvo [former Sigimi], 1886, Jankovski leg. 2♀, Romashka [former Adimi], 1907, Emelyanov leg. 1♀, near Slavyanka , 23.vi.1914, Emelyanov leg. 1♀, Nagornoye , 18.vii.1913, Chersky leg. 9♂, 12♀, Chernigovka , 2 , 30.vi.1912, 11–13, 26.vii.1912, ix.1912, 30.vi.1913, Emelyanov leg. 1♀, Odarka River , 27.vi.1911, Emelyanov leg. 10♂, 8♀, valey of Odarka River, 25 km to Spassk–Dalny [former Evgenyevka] , 26.iv.1911, 26.v.1911, 2, 13, 25, 27, 30.vii.1911, 6, 16.viii.1911, A. Chersky leg. 2♂, Ussuriysk [former Nikolsk–Ussuriysky] , v.1899, Jakovlev leg. 3♀, Ussuriysk , 10.vii.1959, Falkovich leg. 2♀, Ussuriysk [former Nikolsk–Ussuriysky] , 3.vi.1924, 25.vi.1925, Samoy- lova leg. 2♀, Spasskoye , 20 , 17.viii.1910, V. Berger leg. 1♀, Kedrovaya Pad Nature Reserve , 15.vi.1926, Rostovykh leg. 1♂, 2♀, Frolovka , 3, 4 , 6.vii.1926, Rostovykh leg. 1♀, Partizansk [former Suchansky Rudnik], 1934, Pamikov leg. 1♂, Partizansk District [former Suchan District] , 16.vii.1928, Rostovykh leg. 1♂, Partizansk District [former Suchan District] , 3.ix.1949, Borchenius leg. 1♀, Partizansk District [former Suchan District] , 2.vii.1930, R. Malaise leg. 28♂, 33♀, Vinogradovka , 17, 22 , 31.v.1929, 4, 6, 9, 14.vi.1929, 6, 9, 10, 17, 31.vii.1929, 2, 11.viii.1929, Kirit- shenko leg. 4♂, 1♀, Komarovka River [former Suputinka], Reserve , 15–16, 21, 25.vi.1940, 2, 12–13.vii.1940, Ivanov leg. Komarovka River [former Suputinka] , 14.vii.1938, 2♀, Samoylov leg. 2♀, Komarovka River [former Suputinka] , 27.vi.1959, Falkovich leg. 1♂, 3♀, Middle flow of Komarovka River [former Suputinka], tributary of Razdolnaya River [former Suifun] , 18.vi.1936, 2.vii.1938, Samoylov leg. 5♀, Komarovka River [former Suputinka] tributary of Razdolnaya River [former Suifun], 5, 9, 12 , 16.vii.1937, 12.viii.1937, Richter leg. 3♂, near Sokolovka, Kievka River [former Sudzukhe River] , 10–20.vi.1959, Urgens leg. 8♂, 10♀, Kievka River [former Sudzukhe River], Lazovski Nature Reserve [former Primorsky Reserve] , 30.vi.1948, Sharov leg. 4♂, Valley of Serebryanka River [former Sanchoba (Sakhambe, Sanke, Sachenba)], 2, 10 , 23.vi.1945, 5.vii.1945, Nesmerchuk leg. 1♂, NO slope Sikhote-Alin, Samarga River , 17.viii.1932, Prisiezhenkov leg. 1♀, 1 nymph 5 th instar, Gornotaezhnoye , 13– 19.viii.1949, Borchenius leg. 1♂, near Sokolovka , 20–25.vi.1959, Jurgens leg. 1♂, 1♀, Ovchinnikovo [former Vo- roshilov collective farm] , 16.vii.1956, Kamenkova leg. 1♂, 3♀, Alexey–Nikolsky [former Nikolskoye] , viii.1928, Unknown collector. 2♀, near Barabash , 8.viii.1912, Philippovskiy leg. 1♀, Sukhanovka , 17.vii.1912, Sukhanov leg. 1♀, Vladimiro–Alexandrovskoe , 6.vii.1912, Sukhanov leg. 3♂, 2♀, Noveladimirovka, Chersky and Berger leg. Sakhalin Ter.: 1♂, 1♀, Sakhalin Island , Suprunenko leg.

Additional material examined. UKRAINE: Zhytomyr Prov.: 1♂, Zhytomyr Prov. [fomer Volhynian Prov.], 9.vi.1909, A. Kseniopolskiy leg. MONGOLIA: Töv Prov.: 3♂, Noin–Ula, Sutszukte , 28.vi–18.vii.1924, Kozlov leg. 1♂, Sugnugur Gol River [fomer Sugunur], upper portions of the Kharaa River [Kharaa Gol River], 28.vi– 6.vii.1924, Kozlov leg. Dornod Prov.: 1♀, Near Salkhit uul Mount [Numregin–gol], 24.vii.1971, G. Medvedev leg. 1♂, the Khalkh River [The Khalkh–Gol River], 33 km SE to sum [district] Khalkh–Gol , 19.vii.1971, Kerzhner leg. CHINA: Heilongjiang Prov.: 1♂, [Mulin railway station, Chinese Eastern Railway ], 20.vii.1911, A. Emelyanov leg. 1♀, [Imanpo railway station, Chinese Eastern Railway ], 11.vi.1911, 1 nymph 5 th instar, 14.vii.1914, A. Emelyanov leg. 1♀, [Khandaokhetszi railway station, Chinese Eastern Railway, Manchuria ], 15.vii–30.ix. 1918–1921, [coll. Society for the Study of the Manchurian region ] Unknown collector . 1♂, [Gaolintszi railway station, Chinese Eastern Railway, Manchuria ], 3–6.vii.1926, [Society for the Study of the Manchurian region ]. Nanjing Prov .: 1♀, [m. Niu–Shou–Shan, 25 km. Nanking ], 24.vi.1934, Zhenzhurist leg. [identified by Kiritshenko as “ Eurygaster sinicus ”] . 1♀, [ Nanking ], 23.vi.1936, Zhenzhurist leg. Gansu Prov.: 1♂, [ Kansu nur], [identified by Kiritshenko as “ Eurygaster sinicus ”] . 1♀, [ Gan–su ], viii.1872, Przhevalsky leg. 2♀, [near Vey–Yan–Syan], 14.iii.1909, P.K. Kozlov leg. NORTH KOREA: Chagang Prov.: 2♂, 10♀, Coast of the Yalu River , v–vi 1897, V. Komarov leg. [ KOREA]: 1♀, [ Malmi–Sombau ], 18.ix.1900, Schmidt leg. 1♂, [ Kvonkzhen–Syasulma ], 27.ix.1900, Schmidt leg. [ Tekhenge–Kvachen ] , 1♀, 30.ix.1900, Schmidt leg.

I examined the photogaphs of holotype of Eurygaster sinicus Walker, 1867 , sent by Ken Merrifield and Mick Webb (the Natural History Museum, London): CHINA: Hong Kong: Hong Kong, 1♂. I also examined the pho- togaphs of Chinise material sent by Laurent Faivre (the Museum National d’Histoire Naturelle, Paris): Jiangxi Prov .: 1♂, 1♀, [Kiang–si, 1869], A. David leg. 1♂, [Kiang–si, 1875], A. David leg. Guizhou Prov. : 1♀, [Kouy– Tchéou, Rég. De Pin–Fa], 1908, P. Cavalerie leg. Tibet Autonomous Region : 1♂, 2♀, [Thibet (Mou–Pin), 1869– 70], A. David leg.

Diagnosis. Body length 9–11 mm.Apex of clypeus laying clearly below of apices of mandibular plates (anterior view, Fig. 18 ). Apices of mandibular plates twisted clearly inward and in dorsal view ( Fig. 25 ) apices of mandibular plates appear to extend beyond apex of clypeus (in fact, their length is almost the same). Buccula 1.3–1.5 times as high as thickness of second segment of labium. Height of eye 4.7–5.4 times the height of buccula at middle ( Fig. 31 ). Aedeagus with two pairs of conjunctival appendages ( Figs. 37 , 44–49 A–C); appendages of basal pair straight and long; appendages of distal pair shorter and strongly bent almost at right angle in dorsal direction. Blade of crown of paramere ⅓ as long as stem of paramere in dorsal view. Stem at middle distinctly wider than base of crown in dorsal view ( Figs. 44–49 D–F). Laterotergite IX 3.0 or more times as wide as long, usually 0.8-1.0 times as long as valvifer VIII and not reaching lateral excisions of abdominal sternite VII ( Fig. 55 ).

The apices of mandibular plates of all Russian Eurygaster , except E. testudinaria , are almost flat and not twisted inside. Eurygaster testudinaria is very similar to E. maura in the body size and the shape of lateral margins of the pronotum. Eurygaster testudinaria differs from E. dilaticollis , E. integriceps , and E. maura in the apex of the clypeus located clearly below mandibular plates (anterior view, Figs. 13, 15, 17, 18 ). Apices of mandibular plates of E. austriaca and E. laeviuscula are longer than clypeus, and they often converge in front of clypeus. Eurygaster testudinaria is smaller than E. austriaca but generally larger than E. laeviuscula . The anaterolateral margins of pronotum of E. laeviuscula is more convex than in E. testudinaria ( Figs. 23, 25 ).

Natural History and Bionomics. E. testudinaria is a hygro-mesophilic species. It inhabits mostly humid biotopes such as sphagnum bogs overgrown with Eriophorum spp., the wetlands, the flooded oak forests. It is widely oligophytophagous and feeds on Poaceae ( Phleum pretense L., Bromus inermis Leyss. , Festuca spp., Dactylis glomerata L., Anthoxanthum odoratum L., Cynosurus cristatus L., Briza media L., and other species). Adults are sometimes collected on Cyperaceae ( Eriophorum spp., Scirpus spp., Carex spp.), Asteraceae ( Cirsium heterophyllum (L.) Hill, Achillea spp.), Polygonaceae ( Polygonum spp., Rumex spp.), Rosaceae ( Agrimonia spp.), Apiaceae ( Torilis spp.) ( Putchkov 1961). There is one generation per year. The hibernating stage is the imago.

Economic importance. E. testudinaria has not been reported as a pest. In the south of the Russian Far East, this species can be found on cultivated cereals ( Mishchenko 1957, Kanyukova 1995).

Distribution. The species is widely distributed in the Palearctic region and known from North Africa (northern Morocco and Tunisia), almost all of Europe (except the northern and northwestern areas of the Scandinavian Peninsula (Norway), north of Great Britain and most of Ireland), and spanning eastwards from Sinai through Georgia to Japan ( Josifov & Kerzhner 1978, Vinokurov et al. 1988, Vinokurov & Kanyukova 1995, Göllner-Scheiding 2006, Babichev & Vinokurov 2011, Syromyatnikov et al. 2017). In Russia, the species is found almost everywhere in the European Russia and South Siberia and reaching Sakhalin Island in the East; the northern border of distribution passes through the south of Republic of Karelia, the south of Arkhangelsk Province (Kotlas), through the Perm Territory and Sverdlovsk Province, the south of Tyumen (Gornoslinkino) and Tomsk (Semiluzhki) regions, the south of Krasnoyarsk Territory (Tukhtat and Lesosibirsk), through the Irkutsk region (Bunbui and Kirensk), the south of Sakha Republic (Ulahan–Botuobuya River), Khabarovsk Territory (Verkhnebureinsky District), and Sakhalin Island (the first indication from this territory).

Taxonomic notes. Josifov & Kerzhner (1978) lowered E. sinica to a subspecies of E. testudinaria . The subspecies E. t. sinica has East Palaearctic distribution and differs from the nominative subspecies by a shorter distal pair of conjunctival appendages of the aedeagus. I have studied the extensive material from the Zoological Institute of the Russian Academy of Sciences including specimens mentioned by Josifov & Kerzhner (1978). I did not find any difference in the structure of the aedeagus in specimens from the European part and the Far East of Russia as compared to Korean specimens, including the ratio of the lengths of the distal pair of the conjunctival appendages (they are equvalent to “Distalpare Spikula” of Josifov & Kerzhner (1978)). At the same time, I observed the variability in the length of the distal pair of conjunctival appendages of the aedeagus for specimens within the same region for both in the European part and in the Far East ( Figs. 43–48 ). Gaffour-Bensebbane (1992), analyzing and comparing hundreds of male and female specimens of E. testudinaria and E. sinica , also came to the conclusion that these taxa have a complete similarity in the structure of the male and female genitalia, and considered them synonymous. Comparing photos of the aedeagus of E. testudinaria from Russia and adjacent countries and the photos of the aedeagus of the holotype I did not find any differences between them ( Figs. 44–49 , 109–111 ). The lengths of conjunctival appendages of the distal pair are very variable and cannot serve as a reliable criterion for distinguishing these subspecies.

The second feature indicated by Josifov & Kerzhner (1978) for distinguishing of E. t. sinica and E. t. testudinaria is the absence of callouses at the base of the scutellum in E. t. sinica . However, in the collection of the Zoological Institute of the Russian Academy of Sciences there are specimens from Eastern Europe, Siberia, and Mongolia with practically non-prominent callouses. At the same time, I could note that specimens with unclear callouses are much more common in Primorsky Territory of Russia and in Mongolia. On the other hand, there are specimens with a distinct contrasting pattern and clear callouses at the base of scutellum among materials from North Korea, which are mentioned by Josifov & Kerzhner (1978) ( Figs. 86–97 ). The fact that callouses are expressed clearly in E. sinica was mentioned by Péneau (1911). The specimens observed by Péneau are in Figs. 98–103 . In Fig. 108 ( E. sinicus , holotype) the callouses are clearly visible. Thus, I conclude that the presence or absence of callouses is an unreliable feature to divide the species into subspecies E. t. sinica and E. t. testudinaria .

I have studied a part of the material indicated by Josifov & Kerzhner (1978), as well as two females from Nanjing [Nanking] and one male from Gansu [Kansu nur], and did not find any differences between these specimens and specimens from other regions of Russia and Mongolia, which belong to the subspecies E. t. testudinaria ( Göllner-Scheiding 2006) . In my opinion, the material indicated by Josifov & Kerzhner (1978) refers to the nominative subspecies E. t. testudinaria .

The specimens from the collection of the Zoological Institute of the Russian Academy of Sciences mentioned by Josifov & Kerzhner (1978) is stored in a box labeled “ Eurygaster testudinaria ”. According to Nikolay Simov (personal correspondence) the material from the National Museum of Natural History (Sofia), which was examined in this article, is also labeled as “ Eurygaster testudinaria ”.

Gaffour-Bensebbane (1992) considered that E. sinica Walker and E. testudinaria (Geoffroy) are the same species, but subdivided it into two forms: forma sinica , which corresponds to the more massive “ island form ” sensu China (1927) and forma testudinaria , which corresponds to smaller specimens from the continent—“co ntinental form ” sensu China (1927). Studying the material from Great Britain, China (1927) noted that E. testudinaria is larger than E. maura , whereas the opposite situation occurs in the material from the continent. He suggested the existence of two forms of E. testudinaria— island and continental. Thus, Gaffour-Bensebbane (1992) actually equates E. testudinaria from the United Kingdom to E. sinica sensu Péneau (1911) and Walker (1867), who indicated E. sinica from China. This view of Gaffour-Bensebbane (1992) contradicts the accepted division of E. testudinaria into subspecies E. t. testudinaria and E. t. sinica applied in the Catalog of the Heteroptera of the Palaearctic Region ( Göllner-Scheiding 2006). Thus, the size of the body is very unreliable for the division into subspecies. The body size indicated by Walker (1867) and Péneau (1911) are 4 lines and 11 millimetres, respectively. Real length of the holotype body is 9 mm ( Fig. 108 ). This is within my measurements for nominative subspecies of E. testudinaria ( Tab. 1 ).

Gaffour-Bensebbane (1992) also listed other features for separation of these two forms: in E. sinica , the mandibular plates do not cover the clypeus from the sides as much as in E. testudinaria ; in f. sinica the humeral angles of the pronotum extend further beyond the margin of corium than in f. testudinaria ; at the same time, the humeral angles of the pronotum in f. testudinaria are more rounded than in E. maura ; the last three antennal segments are black in f. testudinaria , and can have different color in f. sinica (the shape and length of the antennal segments of both forms are the same); the apices of the middle genital plate of the female in f. sinica are thickened and directed backwards, whereas in f. testudinaria they are evenly rounded and directed forward (in both forms, the median genital plates never reach the margin of sternite VII). According to my observations, all these features are variable in E. testudinaria . In the collection of the Zoological Institute of the Russian Academy of Sciences there are specimens from the European part of Russia and from the Russian Far East with varying levels of covering of the clypeus by the mandibular plates. Additionally, the humeral angles of the pronotum can be more or less rounded and extended beyond the edge of the corium. The colour of the last three antennal segments in E. testudinaria from different regions of Russia can vary from black to brown but the apical segment is more often black.

Thus, in my opinion, it is impossible to identify the features by which one could clearly distinguish the subspecies E. t. sinica from E. t. testudinaria . Based on the foregoing discussion, E. t. sinica Walker, 1867 is synonymized with E. t. testudinaria (Geoffroy, 1785) .

Figures

FIGURES 13–18. Head of Eurygaster, anterior view. 13—E. austriaca (Schrank, 1776), male; 14—E. dilaticollis Dohrn, 1860, female; 15—E. integriceps Puton, 1881, male; 16—E. laeviuscula Jakovlev, 1886, male; 17—E. maura (Linnaeus, 1758), male; 18—E. testudinaria (Geoffroy, 1785), female.

FIGURES 19–25. Head of Eurygaster, view from above. 19—E. austriaca (Schrank, 1776), male (apices of mandibular plates converge); 20—E. austriaca (Schrank, 1776), male (apices of mandibular plates don’t converge); 21—E. dilaticollis Dohrn, 1860, female; 22—E. integriceps Puton, 1881, male; 23—E. laeviuscula Jakovlev, 1886, male; 24—E. maura (Linnaeus, 1758), male; 25—E. testudinaria (Geoffroy, 1785), female.

FIGURES 26–31. Head of Eurygaster, left lateral view (A—photo; B—scheme). 26—E. austriaca (Schrank, 1776), female; 27—E. dilaticollis Dohrn, 1860, male; 28—E. integriceps Puton, 1881, male; 29—E. laeviuscula Jakovlev, 1886, female; 30—E. maura (Linnaeus, 1758), male; 31—E. testudinaria (Geoffroy, 1785), male.

FIGURES 32–37. Schemes of aedeagus of Eurygaster (A—dorsal view, B—dorsolateral view, C—left lateral view). 32—E. austriaca (Schrank, 1776); 33—E. dilaticollis Dohrn, 1860; 34—E. integriceps Puton, 1881; 35—E. laeviuscula Jakovlev, 1886; 36—E. maura (Linnaeus, 1758); 37—E. testudinaria (Geoffroy, 1785).

FIGURES 42–45. Male genitalia of Eurygaster (aedeagus: A—dorsal view, B—dorsolateral view, C—left lateral view; parameres: D—right paramere, E—left paramere, F—left paramere, dorsal view). 42—E. laeviuscula Jakovlev, 1886 (holotype); 43—E. maura (Linnaeus, 1758); 44, 45—E. testudinaria (Geoffroy, 1785).

FIGURES 46–49. Male genitalia of Eurygaster testudinaria (Geoffroy, 1785) (aedeagus: A—dorsal view, B—dorsolateral view, C—left lateral view; parameres: D—right paramere, E—left paramere, F—left paramere, dorsal view). 46—Ukraine, Zhytomyr Province; 47—China, Gansu Province [Kansu nur]; 48—North Korea, Chagang Province; 49—Korea, [Kvonkzhen– Syasulma].

FIGURES 50–55. Female’s postgenital segments of Eurygaster. 50—E. austriaca (Schrank, 1776); 51—E. dilaticollis Dohrn, 1860; 52—E. integriceps Puton, 1881; 53—E. laeviuscula Jakovlev, 1886; 54—E. maura (Linnaeus, 1758); 55—E. testudinaria (Geoffroy, 1785).

FIGURES 86–97. Eurygaster testudinaria (Geoffroy, 1785), variability of coloration. 86, 87—female, Russia, Krasnodar Territory; 88—male, Russia, Kostroma Province; 89—male, Russia, Irkutsk Province; 90—female, Russia, Irkutsk Province; 91—male, Ukraine, Zhytomyr Province; 92—male, Mongolia, Dornod Province; 93—male, China, Nanjing Province; 94, 95, 97—female, North Korea, Chagang Province; 96—male, North Korea, Chagang Province.

FIGURES 98–104. Eurygaster testudinaria (Geoffroy, 1785) from China. 98, 99—male, Jiangxi Province; 100—female, Jiangxi Province; 101, 103—female, Tibet Autonomous Region; 102—male, Tibet Autonomous Region; 104—female, Guizhou Province. © Photo by Laurent Fauvre.

FIGURES 105–111. Eurygaster sinicus Walker, 1867. 105—Holotype labels; 106—anterior view; 107—head, dorsal view; 108—body, dorsal view; 109–111—aedeagus: 109—dorsal view, 110—dorsolateral view, 111—left lateral view. © The Trustees of the Natural History Museum, London. Photography by Ken Merrifield and Mick Webb.