Nannaria stellapolis, Means & Hennen & Marek, 2021

Means, Jackson C., Hennen, Derek A. & Marek, Paul E., 2021, A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae), ZooKeys 1030, pp. 1-180: 1

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scientific name

Nannaria stellapolis

sp. nov.

Nannaria stellapolis   sp. nov. Figs 40 View Figure 40 , 41 Vernacular name: "The Star City Twisted-Claw Millipede" View Figure 41

Material examined.

Holotype: United States - Virginia • ♂; Roanoke County, Roanoke Mountain picnic area off Blue Ridge Parkway ; 37.2309°N, - 79.9502°W; elev. 449 m; 25 Oct.2014; J. Means leg.; VTEC MPE00252 View Materials . GoogleMaps  

Paratype: United States - Virginia • 1 ♂; same collection data as holotype; VMNH MPE00258 View Materials . For detailed collection data see Suppl. material 7 GoogleMaps   .


Adult males of Nannaria stellapolis   sp. nov. are distinct from other Nannaria   and the nearby N. wilsoni   , based on the following combination of characters: Gonopods. Gonopodal acropodite long and curving medially before apex, not ventromedially as in N. hokie   . Acropodite tip with small, triangular lateral flange (Fig. 40A View Figure 40 , red arrow), not encircling tip forming a hood-like structure as in N. hokie   . Acropodite simple, lacking triangular medial flange of N. hokie   . Acropodite without heavily sclerotized laminate corkscrew as in N. wilsoni   . Telopodite basal zone height ca. 1/2 length of acropodite, not < 1/3 length as in N. hokie   or ca. 1/6 length as in N. wilsoni   . Telopodite with small basal zone lateral bulge. Prefemoral process small, thin, curving medially and arising from top of large, projected, shelf-like prefemoral spine-not arising from sharp, acuminate prefemoral spine as in N. hokie   , and not subequal to length of acropodite as in N. wilsoni   . Color. Tergites with orange/red paranotal spots (Fig. 41 View Figure 41 ). Dark brown background. Dorsum of collum smooth with orange anterior margin.


♂ holotype (VTEC, MPE00252): BL = 24.2, CW = 3.3, IW = 1.7, ISW = 0.7, B11W = 3.7, B11H = 2.3.


No known variation.


Known only from the type locality (Virginia: Roanoke County, Suppl. material 7; Fig. 126 View Figure 126 ). Distribution area: N/A; status: MRE.


Individuals of N. stellapolis   sp. nov. were found in mesic broadleaved forests composed of oak, walnut, and maple, under 1-2 cm of hard packed soil on the bank of a dry creek bed.


This species is named for Star City, the common nickname of Roanoke city. The specific name is a noun in apposition derived from the Latin stella -, star, and Greek polis, city.

Type locality.

United States, Virginia, Roanoke County, Roanoke Mountain picnic area off Blue Ridge Parkway, 37.2309°N, - 79.9502°W.