Atholus pirithous (Marseul, 1873)

dela Cruz, Ian Niel & Ohara, Masahiro, 2023, Revision of the genus Atholus Thomson, 1859 (Coleoptera, Histeridae, Histerinae) from the Philippines with additional records, ZooKeys 1158, pp. 1-26 : 1

Atholus pirithous (Marseul, 1873)

Figs 4 , 44-49

Hister pirithous Marseul, 1873: 224 [Japan: Hiogo and Nangasaki].

Hister (Atholus) pirithous : Bickhardt 1910: 54 [catalogued]; 1913: 173; 1917: 194 [catalogued]; Desbordes 1919: 400 [Tonkin]; 1921: 10; Kamiya and Takagi 1938: 31 [listed]; Ôsawa and Nakane 1951: 7.

Atholus pirithous : Lewis 1906, 402; 1915, 55 [Formosa = Taiwan]; Nakane 1981: 10; Mazur 1984: 215; 1997: 132; 2009: 115; 2011: 106 [catalogued]; Mazur et al. 2014: 1269.

Atholus (Euatholus) pirithous : Kryzhanovskij and Reichardt 1976: 390; Hisamatsu and Kusui 1984: 23; Hisamatsu 1985: 223, pl. 41, fig. 19 [key; noted; image]; Ôhara 1993: 141-147; 1994: 138; 1999: 110 [Japan]; 1999b: 36 [Taiwan].

Hister reitteri Bickhardt, 1918: 231 [Japan]; synonymized by Reichardt 1930: 48; Kamiya and Takagi 1938: 31 [listed].

Hister pirithous ab. reitteri : Reichardt 1930: 48.

Specimens examined.

Seven specimens of undetermined sex. Luzon Island , Laguna , northern Lucena, Kinabuhayan, 7 exs. [IC-21-47], 1994-V-VI, N. Monreal leg.

Diagnosis.

Atholus pirithous (Marseul, 1873) is generally recognized for its light excavation in the area behind the anterolateral angle of the pronotum.

Distribution.

Japan, Russia: Far East, China (Guandong, Shanghai), Korea, Taiwan, Vietnam, Nepal, Oman ( Mazur 2011); Philippines (new record).

Biology.

Unknown.

Remarks.

All seven examined individuals of Atholus pirithous (Marseul, 1873) (Fig. 4 ) lack internal subhumeral stria, but traces of dots and short lines can be observed in the apical end. The outer apical protibial tooth of this species is moderately prominent, topped by three denticles. The total number of protibial denticles on the outer margin is ten, one on the inner apical angle (Figs 48 , 49 ), compared with the Japanese specimens described that bore only nine denticles ( Ôhara, 1993), but the outer sublateral margin of the Philippine species has only four denticles; when compared to Ôhara (1993) who observed five to six denticles. Moreover, all examined specimens lost their genitalia prior to examination.

Figures

Figures 1 - 6. Philippine Atholus, dorsal habitus 1 A. philippinensis (Marseul, 1854) [IC- 21 - 18] 2 A. coelestis (Marseul, 1857) [IC- 21 - 20] 3 A. torquatus (Marseul, 1854) [IC- 21 - 49] 4 A. pirithous (Marseul, 1873) [IC- 21 - 47] 5 A. nitidissimus Desbordes, 1925 [syntype image] 6 A. bakeri (Bickhardt, 1914) [syntype image, No. 1639]. Scale bar: 1.00 mm.

Figures 44 - 49. Atholus pirithous (Marseul, 1873) [IC- 21 - 47] 44 habitus, dorsal view 45 ditto, ventral view 46 head, dorsal view 47 propygidium and pygidium 48 protibia, dorsal view 49 ditto, ventral view.