Navira, Piacentini, Luis N. & Grismado, Cristian J., 2009

Piacentini, Luis N. & Grismado, Cristian J., 2009, Lobizon and Navira, two new genera of wolf spiders from Argentina (Araneae: Lycosidae), Zootaxa 2195, pp. 1-33 : 24-27

Navira gen. nov.

Type species. Navira naguan sp. nov.

Etymology. The generic name means “cacique” (tribal leader) in the language of the Henîa people (northern Comechingones), a native extinct aboriginal ethnic group from central Argentina. Gender feminine.

Diagnosis. Navira gen. nov. is clearly distinguished from Lobizon gen. nov. by its glabrous integuments and the unusual transverse banded abdominal pattern, similar among wolf spiders only to the Australian lycosine Mainosa ( Framenau 2006c) . Both characters are here considered apomophies. The palp conformation is very similar to Lobizon gen. nov., but lacks the retrolateral patch of setae on the femur ( Fig. 80 ), the bulb is reduced in relation to the whole palp; the embolus, the median apophysis and the retrolateral tegular lobe are shorter than in Lobizon gen. nov. species ( Figs 77–78 ). Navira gen. nov. can be further distinguished from Lobizon gen. nov. by having three teeth on the cheliceral retromargin ( Fig. 82 ), and by the simpler female genitalia, in which the vulval chambers are inconspicuous (an additional potential apomorphy) and the copulatory ducts are very short ( Figs 13–14 , 75–76 ).

Description. Small wolf spider (TL 4.40–5.93). Females larger than males.

Dorsal shield of prosoma brown, with dark grey radial pattern and light brown marginal bands. Eyes surrounded by black area. Dorsal shield of prosoma without bristles; with four long bristles on the extremely protuding clypeal condyle, anteriorly of AER ( Fig. 73 ). Three promarginal teeth, the median largest; three retromarginal teeth, similar in size. Opisthosoma dark with a light yellow lanceolate mark on the cardiac area and a pattern of pale transverse bands ( Fig. 70 ). Carapace longer than wide, dorsal line straight in lateral view ( Fig. 74 ). Caput flanks in frontal view steep in males, but a gentle slope in females. Leg formula IV> I> III> II. Spination of leg I: femur, two dorsal; tibia, four ventral pairs, metatarsus with three ventral pairs, and one apico-dorsal.

Femur of palp without patch of bristles ( Fig. 80 ), tegulum divided, embolus short, the median apophysis is laminar and reaches the apical border of the tegulum; the terminal apophysis is long, with a curved apical part.

The epigyne is a simple plate with a transverse septum, with the copulatory opening on the lateral borders of the septum; the vulval chambers are inconspicuous and the copulatory ducts are very short. Composition. Only the type species, N. naguan sp. nov.

Figures

FIGURES 79 – 82. 79, 81, Lobizon humilis (Mello-Leitão) male from Reserva Natural Otamendi (MACN-Ar 12514), 80, 82, Navira naguan sp. nov., 80, male without locality (MACN-Ar 16286), 82, female without locality (MACN-Ar 16283). 79, 80, left palpal femur, ventral view. 81, 82, cheliceral retromargin, posterior view.

FIGURES 75 – 78. Navira naguan sp. nov. 75, female paratype, epigyne, ventral (MACN-Ar 16280); 76, female paratype Merlo (MACN-Ar 16280), vulva, cleared, dorsal; 77 – 78, male holotype (MACN-Ar 16281); 77, left palp, retrolateral, 78, same, ventral.

FIGURES 9 – 14. Female genitalia of Lobizon gen. nov. and Navira gen. nov .. 9 – 10, L. corondaensis (Mello-Leitão) from Parque Nacional Chaco, Laguna Carpincho (MACN-Ar 13618); 11 – 12, L. humilis (Mello-Leitão) from Reserva Natural Otamendi (MACN-Ar 16240); 13 – 14, Navira naguan sp. nov. from Bosque Alegre, 20 km de Carlos Paz (MACN-Ar 16282); 9, 11, 13, epigynes, ventral view; 10, 12, 14, vulvae, dorsal view. Upper right corner in Fig. 12, detail of the dictynoid pore; upper right corner in Fig. 14, detail of the pores of the head of right spermatheca.

FIGURES 69 – 74. Navira naguan sp. nov., habitus of preserved specimens from Merlo. 69, 71, female paratype (MACN-Ar 16280); 70, 72 – 74, male holotype (MACN-Ar 16281); 69 – 70, dorsal; 71 – 72, ventral; 74, lateral; 73, detail of the ocular area and the clypeal condyle.