Coptoprepes casablanca, Werenkraut, Victoria & Ramírez, Martín J., 2009

Werenkraut, Victoria & Ramírez, Martín J., 2009, A revision and phylogenetic analysis of the spider genus Coptoprepes Simon (Araneae: Anyphaenidae, Amaurobioidinae), Zootaxa 2212, pp. 1-40: 18-23

publication ID

http://doi.org/ 10.5281/zenodo.275145

persistent identifier

http://treatment.plazi.org/id/C927878D-FFCC-CB4B-5AA6-FA46FED6A0C7

treatment provided by

Plazi

scientific name

Coptoprepes casablanca
status

new species

Coptoprepes casablanca   new species

Figs. 11 –14 View FIGURE 11 View FIGURE 12 View FIGURE 13 View FIGURE 14 , 22 View FIGURE 22 c, 23g –i, 24g –i

Type material: Holotype: male from Chile, Región X (Los Lagos), Osorno province, Puyehue Natl. Park, Volcán Casablanca   , ca. 40 º 30 ’ S, 72 º00’ W, 1130–1180 m, site 665, pan traps above treeline, Valdivian rainforest, 20–25 December 1982, A. Newton & M. Thayer ( AMNH; temporary preparations MJR-01330, VIW-00005). Paratype: female from Chile, Región X (Los Lagos), Puyehue Natl. Park, Antillanca rd., 1035 m, Nothofagus   forest, 31 January 1985, N. I. Platnick & O. F. Francke ( AMNH; temporary preparations MJR-01338, VIW- 00006).

Etymology: The specific name is a noun in apposition taken from the type locality.

Diagnosis: Females are similar to that of C. eden   n. sp. by having the lateral lobes of the epigyne posteriorly concave ( Fig. 13 View FIGURE 13 c), continued in a pouch leading to the copulatory openings, but can be distinguished by the larger, almost spherical spermathecae ( Fig. 24 View FIGURE 24 g –i). Males resemble those of C. flavopilosus   by having a curved RTA and relatively short embolus, with the tegulum visible in ventral view ( Fig. 13 View FIGURE 13 b –d), but differ by having simple, hook-shaped, slender median apophysis ( Fig. 23 View FIGURE 23 g –i).

Description: Male (holotype): Prosoma length 2.10, width 1.37, narrower in front than in the female. Tibia/metatarsus length: I, 1.17 / 1.12; II, 1.02 / 0.97; III, 0.82 / 1.02; IV, 1.17 / 1.42. Chelicerae slightly smaller than those of female, with five teeth on retromargin, five on promargin. Sternum length 1.07, width 0.85. Spines: leg I, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 - 2; metatarsus v 2 bas. II, femur d 1 - 1 - 1, p d 1 ap; tibia r 1 -r 1 -p 1 or r 1 -r 1-2; metatarsus v 2 bas. III, femur d 1 - 1 - 1, p 0-d 1 -d 1, r d 1 ap; patella r d 1; tibia v p 1-2 - 2, p 1 -d 1 - 1 -0, r d 1 - 1, d r 1 bas; metatarsus v 2 -p 1 -0 plus apical comb, p and r d 1 - 1 - 1, d 0-p 1- 2. IV, femur d 1 - 1 - 1, p and r d 1 ap; patella r d 1; tibia v p 1-2 - 2, p 1 -d 1 - 1 -0, r 1 -d 1 -0-0, d r 1 bas; metatarsus v 2 - 2 -0 plus apical comb, p and r d 1 - 1 - 1, d 0-2 - 2. Opisthosoma length 2.43, spiracle –epigastrium 1.32, spiracle –spinnerets 0.20. Color in ethanol ( Fig. 11 View FIGURE 11 ): As in female but slightly decolorated, the patches of white setae on abdomen on pale cuticle. Palp ( Fig. 13 View FIGURE 13 ): Tibia short, width/length 1.33, RTA shorter than tibia, compressed, curved. Cymbial conductor wide. Tegulum displaced basally in ventral view. Sperm duct with conspicuous loop on anterior-dorsal margin. Embolus thin, with flat rounded basal process. Median apophysis with large base and long, acute, hook-shaped tip. Primary conductor well sclerotized, with canal, ending in acute tip. Secondary conductor a broad concave ledge forming a canal, arising from sclerotized area continuous with anterior-dorsal margin of tegulum, dorsal side regularly covered with denticles not reaching the tip. Paramedian apophysis absent ( Fig. 23 View FIGURE 23 g –i).

Female (paratype): Prosoma length 2.27, width 0.82. Length of tibia/metatarsus: I, 1.00/ 0.92; II, 0.98 / 0.87; III, 0.80 / 1.02, 1.20 / 1.50. Chelicerae with five (right) or six (left) teeth on retromargin, five on promargin. Sternum length 1.17, width 0.90. Spines: leg I, femur d 1 - 1 - 1, p 2 ap; tibia v 2 - 2 -p 1 or 2 - 2 -0; metatarsus v 2 bas. II, femur d 1 - 1 - 1, p d 1 ap; tibia v r 1 -r 1 -p 1; metatarsus v 2 bas. III, femur d 1 - 1 - 1, p 0-d 1 -d 1, r d 1 ap; patella r d 1; tibia v p 1 -p 1-2, p 1 -d 1 - 1 -0, r d 1 - 1, d r 1 bas; metatarsus v 2 - 2 -0 plus apical comb, p and r d 1 - 1 - 1, d 0-2 - 2. IV, femur d 1 - 1 - 1, p and r d 1 ap; patella r d 1; tibia v p 1-2 - 2, p and r 1 -d 1 - 1 -0, d r 1 bas; metatarsus v 2 - 2 -0 plus apical comb, p and r d 1 - 1 - 1, d 0-2 - 2. Color in ethanol ( Fig. 12 View FIGURE 12 ): Prosoma and legs dark brown, femora with paler ventral distal spots, patellae and tibiae with longitudinal paler bands. Abdomen dark brown, with paler muscle insertions, two pairs of dorsal lateral patches of white hairs; venter paler, with four longitudinal lines of pale spots. Opisthosoma length 3.23, epigastrium –spiracle 1.55, spiracle –spinnerets 0.33. Epigyne: Lateral lobes separated from median field, concave, continued in pouch to copulatory openings, close to epigastric furrow. Median field weakly sclerotized, slightly elevated ( Fig. 13 View FIGURE 13 g). Copulatory ducts not wrapped. Ducts of accessory bulbs short. Spermathecae near spherical, with ample lumen ( Figs. 13 View FIGURE 13 h, 24g –i).

Variability: The female internal genitalia have been found to be quite variable, often asymmetric ( Fig. 14 View FIGURE 14 ). This may be related with being abnormally developed, as specimens often had scars on the abdominal cuticle compatible with those produced by nematode parasitoids ( Fig. 14 View FIGURE 14 a). Spines in males: III, metatarsusus, d 0-2 - 2.

Distribution: Known for Río Negro province in Argentina, and for Cautín, Malleco, Osorno provinces, and one isolated record in Aisén province in Chile, which suggest that it may occur in intermediate localities in Palena province.

Other material examined: ARGENTINA: Río Negro: Cerro Chall-huaco, S 41.25718 º, W 71.28316 º, 1300 m, pitfall traps (cod. M 5 S 5 E06), January 2006, V. Werenkraut, 1 female (MACN-Ar 19161). CHILE: Región IX (Araucanía): Cautín: Volcán Villarrica, 1250 m, site 653, window trap, Nothofagus dombeyi   - pumilio   forest with Chusquea   , 15–29 December 1982, A. Newton & M. Thayer, 5 males ( AMNH); same data, baited pitfall traps, 3 males ( AMNH); site 654, window trap, Nothofagus dombeyi   / Saxegothea   forest with Drimys   , 15–29 December 1982, A. Newton & M. Thayer, 3 males [plus 1 immature male] ( AMNH). Malleco: 14 km E Malalcahuello, 1570 m, site 649, window trap, Nothofagus pumilio   / Araucaria   forest, 13–31 December 1982, A. Newton & M. Thayer, 1 male ( AMNH); 12 km E Malalcahuello, 1350 m, site 650, window trap, Nothofagus dombeyi   / Araucaria   forest, 13.31 December 1982, A. Newton & M. Thayer, 2 males ( AMNH); 14 km E Malalcahuello, 1525 m, Araucaria   forest, 25 January 1985, N. Platnick & O. Francke, 1 female ( AMNH; temporary preparation MJR-01355). Región X (Los Lagos): Osorno: Puyehue Natl. Park, Volcán Casablanca   , 1270 m, trap site 668, carrion trap (squid), just above tree line, 22–25 December 1982, A. Newton & M. Thayer, 1 male ( AMNH; temporary preparation MJR-01331); 1130–1180 m, site 665, pan traps above treeline, Valdivian rainforest, 20–25 December 1982, A. Newton & M. Thayer, 4 males and 1 female ( AMNH; temporary preparation MJR-01352); 1250 m, tree line, 31 January 1985, N. I. Platnick & O. F. Francke, 1 female ( AMNH; temporary preparation MJR-01350), 2 females ( AMNH; temporary preparations MJR-01351, MJR-01354). Región XI (Ibáñez del Campo): Aisén: Reserva Nac. Cerro Castillo, 500–600 m, dry forest, 7 February 1985, N. I. Platnick & O. F. Francke, 1 female ( AMNH; temporary preparations MJR-01353, VIW-00019).

AMNH

American Museum of Natural History