Reticunassa goliath, Galindo & Kool & Dekker, 2017

Galindo, Lee Ann, Kool, Hugo H. & Dekker, Henk, 2017, Review of the Nassarius pauperus (Gould, 1850) complex (Nassariidae): Part 3, reinstatement of the genus Reticunassa, with the description of six new species, European Journal of Taxonomy 275, pp. 1-43 : 31-34

Reticunassa goliath sp. nov.

urn:lsid:zoobank.org:act:5C34E90F-3AD2-42ED-B7A6-582525AD11DB

Fig. 8 E–H, 10; Table 2

Etymology

This species is named after the biblical Goliath, the giant who fought against David. This name is chosen because Reticunassa goliath sp. nov. is to date the largest known species in the genus Reticunassa .

Type material

Holotype

NEW CALEDONIA: Nouméa , in sandy areas within coral reef, snorkeling at night, leg. D. Massemin ( MNHN IM-2000-22724 , ex coll. H. Kool, length 15.4 mm, width 7.9 mm).

Paratypes

NEW CALEDONIA: PLOUVEAL, stn 1221, Lagon d’Ouvéa, 20°29′ S, 166°31′ E, 10 m ( MNHN IM- 2000-22725, MNHN IM-2000-22723, MNHN IM-2000-28394, 3 spm; ZMA.Moll.4.09.050, 1 spm; GH, 1 spm; HD 24143, 2 spm; HK 997.01, 3 spm; HUJ, 1 spm); PLOUVEAL, stn 1227, 12°37′ S, 166°25′ E, 12 m ( HD 35304, 2 spm; HK 997.05, 2 spm); Nouméa, sand between coral reefs, snorkeled at night ( HK 997.07, 1 spm); Nouméa, Nouville Beach, brown sand, 10 m ( HK 997.08, 1 spm); New Caledonia, 5–10 m ( HD 26939, 1 spm).

Other material examined

INDONESIA: West Papua, Manokwari, leg. D. Smits ( HK 997.03, 2 spm).

AUSTRALIA: North Queensland, Bowen, Grey’s Bay, ex coll. Hessel ( ZMA, 4 spm; HK 997.06, 1 spm).

VANUATA: SANTO 2006, stn FB52, Malokilikili, 15°42.7′ S, 167°15.1′ E, 7 m (1 spm).

NEW CALEDONIA: LAGON: stn 161, Ile Ouen-Baie du Prony, 22°34′ S, 166°38′ E, 20 m (1 spm); stn 943, Koumac, 20°37′ S, 164°11′ E, 15 m (1 spm). – PLOUVEAL, Lagon d’Ouvéa: stn 1218, 20°36′ S, 166°30′ E, 13 m (8 spm); stn 1219, 20°30′ S, 166°28′ E, 15 m (26 spm); stn 1220, 20°29′ S, 166°29′ E, 14 m (4 spm); stn 1221, 20°29′ S, 166°31′ E, 10 m (69 spm); stn 1222, 20°28′ S, 166°30′ E, 15 m (13 spm); stn 1223, 20°28′ S, 166°28′ E, 19 m (1 spm); stn 1224, 20°32′ S, 166°28′ E, 18 m (25 spm); stn 1225, 20°36′ S, 166°28′ E, 18 m (6 spm); stn 1226, 20°32′ S, 166°24′ E, 21 m (5 spm); stn 1227, 12°37′ S, 166°25′ E, 12 m (29 spm); stn 1228, 20°36′ S, 166°24′ E, 18 m (8 spm); stn 1229, 20°37′ S, 166°23′ E, 16 m (19 spm); stn 1230, 20°35′ S, 166°23′ E, 18 m (22 spm); stn 1231, 20°31′ S, 166°23′ E, 23 m, 6 spm); stn 1232, 20°32′ S, 166°24′ E, 31 m (7 spm); stn 1233, 20°29.1′ S, 166°29.0′ E, 15 m (1 spm). – MONTROUZIER 1993: Koumac, stn 1271, 20°52.7′ S, 165°19.5′ E, 5–25 m (3 spm); Koumac, stn 1286, Plateau Karembé, 20°38′– 20°39′ S, 164°16′– 164°17′ E (3 spm); Koumac, stn 1287, Récif de l’Infernet, 20°37′ S, 164°14′ E (2 spm); Koumac, stn 1301, Récif de l’Infernet, 20°37.1′– 20°37.5′ S, 164°14.7′– 164°15′ E, 1–5 m (3 spm); Koumac, stn 1303, Lagon, parages du Plateau Karembé, 20°37.7′– 20°38.8′ S, 164°15.9′– 164°17.1′ E, 0–8 m (2 spm); Koumac, stn 1304, Chenal de l’Infernet, 20°38.6′ S, 164°13.2′ E, 12–15 m (2 spm); Koumac, stn 1306, Chenal de l’Infernet, 20°39.1′ S, 164°12.4′ E, 11–13 m (1 spm); Koumac, stn 1312, 20°40′ S, 164°14.9′ E, 26–40 m (1 spm); Touho, stn 1245, Grand Récif Mengalia, 20°45.2′ S, 165°16.3′ E (4 spm); Touho, stn 1273, Touho region, outer reef, 20°50.4′ S, 165°22.8′ E, 20 m (2 spm). – PALEO-SURPRISE 1999, stn CP 1388, 18°23.8′ S, 163°06.9′ E, 40 m (2 spm).

LOYALTY ISLANDS:LIFOU 2000, stn 1419,Lifou, Santal Bay, Bay of Gaatcha, 20°55.6′ S, 167°04.5′E, 5 m (8 spm; HK 997.02, 1 spm); LIFOU 2000, stn 1426, Lifou, Santal Bay, 20°45.9′ S, 167°06.2′ E, 4–7 m (2 spm); Lifou, leg. J.R. le B. Tomlin ( ZMA.Moll.096228, ex coll. Schepman, 6 spm); Lifou, leg. R.P. Goubin ( ZMA.Moll.089599, ex coll. Dautzenberg, 3 spm); Lifou ( HUJ 9055, ex coll. Dautzenberg, ex coll. Coen 7553, 3 spm).

Description

Holotype

PROTOCONCH. Paucispiral, pointed, 1.75 milky whorls, beset with rows of microscopic pustules. Protoconch missing in holotype, seen in paratype ( Fig. 8H ).

SHELL. Rather heavy, conical, acuminate, teleoconch consisting of 7 whorls, suture impressed. On penultimate 13 and on body whorl 12 well pronounced, equidistant axial ribs and strong varix. Ribs equally strong on dorsal and ventral side of body whorl.

SPIRAL CORDS. Continuous, moderately broad on top of axial ribs, but between ribs narrower and weak, approximately 6 cords on penultimate and 10 on body whorl.

INTERCORDAL SCULPTURE. Numerous very fine, evenly spaced spiral striae, occasionally also on top of spiral cords.

APERTURE. Round, proportionally small, approximately 1/5 of shell length. Outer lip with 7 unequal denticles. Columella callused, somewhat elevated anteriorly and with evenly spaced lirae throughout. Callus sharply bordered, bending over part of fasciole and somewhat extending over body whorl posteriorly. Inside outer lip with 8 lirae and pronounced tooth at siphonal canal. Elevated part of callus with fine growth lines on left (out)side. Parietal denticle strong, anal canal deep.

OPERCULUM. Yellowish, serrated.

SIPHONAL CANAL. Narrow, fasciole strong. Siphonal area with one strong and some weak cords.

COLOR. Off white to yellow, narrow band just above suture and broader band below periphery; columella and outer lip white, aperture yellowish.

ADULT SIZE. 8.2–16.5 mm, usually 13–15 mm.

Remarks

Color and banding pattern variable; some specimens have light to dark brown bands and others lack banding. In banded specimens, one band occurs on the body whorl, with an additional band on the sutural area. In some specimens, the spiral cords may be colored orange/brown, but only between the axial ribs. Aperture white, inside faintly showing the outside bands if present.

Reticunassa visayaensis sp. nov. and R. poppeorum sp. nov. have a multispiral protoconch, whereas the protoconch of R. goliath sp. nov. is paucispiral.

The strong columellar callus, together with the thickening of the inside of the outer lip and the strong varix, gives the aperture a round, “open mouth” appearance, whereas the other species in this group have a more oval-shaped aperture.

R. goliath sp. nov. and R. tringa both have a paucispiral protoconch, but the former has a considerably heavier and larger shell. It is similar to R. tringa , R. visayaensis sp. nov. and R. poppeorum sp. nov. in having weak spiral cords. R. goliath sp. nov. differs from R. paupera in weight, in its larger size, in the number of whorls, and in the size of the aperture relative to the total shell length.

Habitat

Sublittoral to 0–40 m, frequently between 10 and 20 m.

Distribution

Western Pacific; Indonesia (Papua), Australia (Queensland), Vanuatu, New Caledonia, and Loyalty Islands ( Fig. 10 ).

Figures

Fig. 8. Shells of Reticunassa Iredale, 1936. – A–D. Reticunassa annabolteae sp. nov. A–C. Holotype, MNHN IM-2009-12843, length 11.5 mm, width5.3 mm, S Madagascar,Ambatobe, Bavarama,25°27.9′ S, 44°57.6′ E, 0–1 m. D. Protoconch. – E–H. R. goliath sp. nov. E–G. Holotype, MNHN IM-2000- 22724, length 15.5 mm, width 7.9 mm, New Caledonia, Nouméa, leg. D. Massemin, ex coll. H. Kool. H. Protoconch of paratype, MNHN IM-2000-28394. – I–L. R. intrudens sp. nov. Holotype, MNHN IM-2000-22726, length 7.9 mm, width 4.3 mm, New Caledonia, Touho, 20°44′ S, 165°14′ E, 49–59 m. L. Protoconch. – M–P. R. thailandensis sp. nov. M–O. Holotype, ZMA Moll.4.09.047, length 7.35 mm, width 3.8 mm, Thailand, Gulf of Thailand, 6°40.313′ N, 101°43.852′ E, 3–15 m. P. Protoconch. Scale bars: D, H, L, P = 200 µm.

Fig. 10. Geographical and bathymetrical distribution of Reticunassa goliath sp. nov. Star indicates type locality. Each bar represents all lv or dd specimens.