Ayacucho bambamarca (Roewer, 1957) Benedetti & Pinto-da-Rocha, 2022

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo, 2022, Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species, Arthropod Systematics & amp; Phylogeny 80, pp. 309-388 : 309

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Ayacucho bambamarca (Roewer, 1957)

comb. nov.

3.11. Ayacucho bambamarca (Roewer, 1957) comb. nov.

Figs 2A View Figure 2 , 7A, B View Figure 7 , 12A, B View Figure 12 , 28 View Figure 28

Cajamarca bambamarca Roewer, 1957: 76 (desc.), fig. 33 (male femur IV); Kury 2003: 144 (cat.).

Cajamarca triseriata Roewer, 1957: 75 (desc.), fig. 32 (male femur IV); Kury 2003: 144 (cat.), syn. n.


MALE: Measurements (n =7) DSW: 3.6-5.1 (5.1); DSL: 3.8-5.2 (5.2); CL: 1.5-1.7 (1.7). FIVL: 4.4-5.2 (4.6). ChL: 1.5-3.7 (3.6). Coloration (in ethanol): Predominantly yellowish. Dark spots on carapace. Areas I-IV, lateral and posterior margin of dorsal scutum and free tergites I-III more brownish. Dorsum: (Fig. 2A View Figure 2 ) Alpha-type DSS. Anterior margin of dorsal scutum with median elevation, granules distributed throughout its length. Ocularium with a pair of spines and few granules. Carapace with scattered granules distributed throughout its length. Areas I-IV densely granulate; area I with a pair of median tubercles slightly larger than the surrounding granules; area II with a pair of small median tubercles, larger than that of area I. Areas III-IV with a pair of small tubercles. Posterior margin of dorsal scutum and free tergites I-III with a row of acuminate tubercles, the median larger than that on areas I-IV. Lateral margins of dorsal scutum with granules distributed throughout their length. Chelicerae: (Fig. 2A View Figure 2 ) Swollen in large males (as in the holotype), similar to females in the small males. Segment I granulate. Segment II predominantly smooth, finger with one tooth. Segment III with two teeth. Pedipalpus : Small granules distributed on the dorsal surface of the femur and patella. Trochanter with a large ventroapical setiferous tubercle. Femur with a ventrobasal setiferous tubercle; a row of six ventral setiferous tubercles, except at base and apex, larger in larger males. Tibia: prolateral III, retrolateral iiIi. Tarsus: prolateral IIi, retrolateral iIiii. Venter: Coxa I with a median row of six small tubercles. Coxae II-IV smooth. With three tiny tubercles apically between coxae II-III and III-IV. Smooth genital area. Free sternites with a row of small granules. Anal operculum with granules sparsely distributed throughout. Legs: (Figs 2A View Figure 2 , 7A, B View Figure 7 ) Coxae II-III with a prolateral apophysis. Coxa IV with granules distributed throughout its surface, and a proapical apophysis with acuminate apex. Trochanters I-III smooth. Trochanter IV with dorsal granules and an apical retrodorsal acuminate tubercle. Femora I-II with granules scattered throughout their length; III densely granulate; with retro and prolateral rows of small acuminate tubercles; IV granulate; with a dorsal row of 7-9 (8) large acuminate tubercle, at to basal ⅔, growing apically; a retroventral row of 16-23 (23) large acuminate tubercles; a retroventral apical tubercle; a proventral row of 10-11 (11) large acuminate tubercles, at ⅔ of its length apically. Patellae I-III with granulation throughout their extension; IV with dorsoapical tubercles. Tibiae I-IV unarmed, with granules throughout their length. Tarsal formula: (n =8) 7, 12-13 (13), 6-7 (7), 7-8. Penis: (Fig. 12A, B View Figure 12 ) VP subrectangular; distal margin slightly concave, with lateral projections; straight in lateral view. MS C1-C3 subapical long, slender and straight or slightly curved; MS A1 and MS B1 short and straight, placed next to MS C. Lateral sacs long and apically acuminate, with long T3-like microsetae. Stylus slightly thick, apically inflated, with tiny apical projections. - FEMALE: Measurements (n =4) DSW: 3.4-4.7; DSL: 3.8-5.0; CL: 1.2-1.8. FIVL: 3.7-4.5. ChL: 1.3-2.3. Chelicerae similar to that of small males. Presence of a proapical spine in femur of pedipalpus. Femora III-IV unarmed. Tarsal segmentation: (n =4) 7, 10-12, 7, 8-9.


Similar to Ayacucho triarmatus nom. nov. and A. weyrauchi comb. nov. by possessing three rows of spiniform tubercles in male femur IV (Fig. 7A, B View Figure 7 ). It differs from A. weyrauchi comb. nov. for having more than four spiniform tubercles in retroventral row of femur IV (Fig. 7B View Figure 7 ); higher tubercles in free tergites (Fig. 2A View Figure 2 ); areas I-IV with a pair of median tubercles (Fig. 2A View Figure 2 ); dorsal process absent in the penis (Fig. 12A, B View Figure 12 ). Differs from A. triarmatus nom. nov. by possessing the retroventral row of spiniform tubercles along entire length of femur IV (Fig. 7B View Figure 7 ); apical margin of the ventral plate with conspicuous lateral projections (Fig. 12A, B View Figure 12 ); dorsal process absent (Fig. 12A, B View Figure 12 ).


Considering that Cajamarca bambamarca and C. triseriata were described in the same work ( Roewer 1957), we established the precedence of C. bambamarca (Art. 24.2 of ICZN). Regarding the type of C. bambamarca : Roewer designated one male as holotype and five males as paratypes, but the type material is preserved without separation in the same vial. Therefore, it is not possible to recognize with absolute certainty which of the males is the holotype. As a consequence of this, one of the males, whose femur IV most closely resembles the drawing in the original description, was separated as the holotype (although it is important to point out that the drawing does not faithfully represent any of the specimens).


(Fig. 28 View Figure 28 ) PERU. Cajamarca . Bambamarca; Cerro Macheipungo.

Material examined.

Type material: Of C. bambamarca : Holotype ♂, ' PERU, Cajamarca , Bambamarca | 2,800m | 29/VI/1956, Weyrauch leg. (SMF RII 11649 View Materials /32) - Paratypes 5 ♂, 3 ♀ ‘ditto’ (SMF RII 11649 View Materials /32). Of C. triseriata : Holotype ♂ ' PERU, Cajamarca , Cerro Macheipungo | 4 km NE Bambamarca, 3,000m | 28/VI/1956, Weyrauch leg. (SMF RII 11647 View Materials /30) - Paratype ♀ ‘ditto’ (SMF RII 11647 View Materials /30) .














Ayacucho bambamarca (Roewer, 1957)

Benedetti, Alipio Rezende & Pinto-da-Rocha, Ricardo 2022

Cajamarca bambamarca

Benedetti & Pinto-da-Rocha 2022

Cajamarca triseriata

Benedetti & Pinto-da-Rocha 2022