Callistomorphus malleus, Mazur, Milosz Adam, 2019

Mazur, Milosz Adam, 2019, Redescription of the forgotten New Caledonian weevil genus Callistomorphus Perroud, 1865 (Coleoptera, Curculionidae, Eugnomini) with descriptions of eight new species, ZooKeys 821, pp. 45-83 : 45

Callistomorphus malleus sp. n. Figs 2, 10, 16, 25, 34, 43, 52, 61, 70, 79, 88, 97, 104, 111, 118, 123, 128, 133, 136, 141

Diagnosis.

This species can be distinguished from other members of the genus by the following suite of characters: rostrum gently curved, regularly narrowed to apex in lateral view. Middle tubercles on elytra flattened, lower than maximal width at base. Apex of penis in lateral view expanded into small tubercles. Apical lobes of female abdominal sternite VIII with characteristic shape (Fig. 136).

Description.

Body length (lb) - 8.60-10.50 mm.

Body colour and vestiture (Fig. 16). Colour variable, from dark brown to yellowish. Some specimens with whitish coating on apical and lateral parts of elytra. Similar, pale coating, in some specimens, also covered hind femora. Pronotum with longitudinal yellowish stripes.

Head (Figs 61, 70, 79). From subquadrate to elongate (hw/hl ♂: 1.00-1.20; ♀: 1.00-1.33), depending on protrusion the head from pronotum. Frons wider than double width of eye. Eyes approximately half as long as head or little longer (eyl/hl ♂: 0.50-0.60; ♀: 0.45-0.55), not protruding above margin of head in lateral view; regularly convex, widest near middle of length. Rostrum longer than pronotum (rl/pl ♂: 1.17-1.21; ♀: 1.12-1.24); in male 3.80-4.30 × as long as maximum width, in female 4.00-4.50 × as long as wide at apex (rl/arw); longitudinal carina indistinct, from base to antennal insertion covered by scales, apically bare, flattened, sometimes evanescent and visible only as polished, narrowed area. Scape shorter than rostrum (scl/rl ♂: 0.75-0.80; ♀: 0.70-0.85). First funicle segment ca 1.3 × as long as 2nd; 3rd almost as half-length of 2nd, antennomeres from 4th to 7th with similar length, slightly longer than wide. Club as long as last four funicle segment combined, approximately 2.10 × as long as wide.

Pronotum (Figs 43, 52). Subquadrate (bpw/pl ♂: 1.00-1.05; ♀: 0.95-1.10). Base in male 1.25-1.35 ×, in female 1.28-1.38 ×, as wide as apical margin (bpw/apw); apical margin straight or slightly concave; in lateral view apical margin straight anteriorly, then converging towards base. Medial tubercle on pronotal disc well developed but weakly protruding. Width of medial constriction in relation to apical and basal margin: mpw/apw ♂: 0.66-0.68; ♀: 0.61-0.69, mpw/bpw ♂: 0.48-0.53; ♀: 0.48-0.50.

Elytra (Figs 25, 34). Elongate (el/bew ♂: 1.56-1.66; ♀: 1.50-1.68); lateral margins subparallel basaly to apical convergence. Medial tubercle distinct but flattened, lower than width on base; smaller, numerous tubercles on elytral disc acuminate; tubercles on 7th intervals clearly visible, protruding from outline of elytra in dorsal view. Striae with small, suboval, shallow punctures; surface of elytral disc slightly glabrous. Scutellum slightly longer than wide.

Abdomen (Figs 88, 104, 128). From slightly (in male) to distinctly (in female) longer than wide (al/aw ♂: 1.08-1.17; ♀: 1.06-1.20 × as long as wide. Male pygidium as in Fig. 104, female as in Fig. 129. Last ventrite longer than wide (lvw/lvl ♂: 2.25-2.57; ♀: 2.13-2.38).

Male terminalia (Figs 97, 111, 118). Penis body slightly longer than apodemes; from base to apical part almost subparallel, apically slightly narrowed to rounded apex; basal part sclerotised, except medial area; in lateral view distinctly, regularly curved, apically expanded into small tubercles. Internal sac without any structure or sclerites. Parameroid lobes of tegmen thin, distinctly shorter than apodeme, divided almost to base and surrounded by thin membrane. Tegminal apodeme apically extended. Spiculum gastrale robust, apodeme laterally flattened, distinctly bent distally; hemisternites fused with base of spiculum.

Female terminalia (Figs 123, 128, 133, 136, 141). Apical lobe of abdominal sternite VIII with characteristic, T-shape sclerotisation. Abdominal tergite VIII with elongate, straight, dense setae near apex; sides with strongly elongate setae; apex with small, pointed tubercles. Spermatheca elongate, strongly bent. Gonocoxite placed diagonally to partly sclerotised vagina, strongly elongate, apically with bundle of erect setae.

Measurements. ♂: al 3.50-4.00, apw, 1.60-1.80, arw 0.60-0.70, aw 3.00-3.7, bew 3.50-4.30, bpw 2.00-2.40, el 5.80-6.90, eyl 0.60, frw 0.40-0.50, hl 1.00-1.20, hw 1.15-1.20, lb 8.60-10.40, lvl 0.70-0.80, lvw 1.60-2.00, mpw 1.05-1.20, pl 1.90-2.40, rl 2.30-2.80, scl 1.80-2.20.

♀: al 3.80-4.10, apw, 1.60-1.80, arw 0.60-0.70, aw 3.30-3.80, bew 3.90-4.30, bpw 2.20-2.40, el 6.00-7.20, eyl 0.50-0.60, frw 0.50-0.60, hl 0.9-1.20, hw 1.10-1.25, lb 9.50-10.50, lvl 0.80, lvw 1.70-2.00, mpw 1.10-1.20, pl 2.10-2.50, rl 2.50-2.80, scl 1.90-2.10.

Type material.

Holotype, ♂ (here designated) - New Caledonia (S); 22°04'08.9"S, 166°26'48.0"E; Dzumac road; S of Mts Couvélé rd jct; 870→670 m beating; 31.10.2008; leg. M. Wanat (MNHN).

Paratypes:

1♂ - New Caledonia (S); 22°05.9'S, 168°38.3'E; Riviére Bleue Parc; 23.12.2006, 190 m, refuge; sifting forest litter; leg. R. Dobosz (USMB).

1♂ - New Caledonia (S); 22°14.9'S, 166°49.7'E; Pic du Pin, base; 25.12.2006, 280 m, forest & plantation; leg. R. Dobosz & M. Wanat (USMB).

1♂ - New Caledonia (S); 22°10'19.2"S, 166°45'40.0"E; Bois du Sud, 220 m, at light; 25.10.2008; leg. M. Wanat (MNHW).

1♀ - New Caledonia (S); 22°10'22.4"S, 166°45'47.9"E; Bois du Sud, 220-250 m, beating along track entering forest reserve; 20.10.2008; leg. M. Wanat (MNHW).

1♀ - New Caledonia (S); 22°05.8'S, 166°40.2'E; Riviére Bleue: Gue de la; 22.12.2006, 140 m Pourina; night coll. (lamp & beating); leg. M. Wanat & R. Dobosz (MNHW).

1♀ - New Caledonia (S); 22°01.9'S, 166°28.0'E; Dzumac Mts, 900 m; Mt. Ouin, road junction; 28.12.2006, night collecting; leg. M. Wanat & R. Dobosz (MNHW).

2♀♀ - New Caledonia (S); 22°12'21.2"S, 166°40'46.9"E; Col des Deux Tétons, 30.10.2010; humid forest, 220-250 m; leg. M. Wanat & R. Ruta (MNHW).

1♀ - New Caledonia (S); 22°06.0'S, 166°39.3'E; Riviére Bleue, Pont Germain to kaori géant (left river side), 160-180 m; 22.01.2007; leg. M. Wanat (MNHW).

Etymology.

This epithet is derived from the Latin noun " malleus " (hammer) and refers to the shape of female sternite VIII. A noun in apposition.

Remarks.

The species is variable in size and colour but easily distinguished by the elongate elytra with relatively small medial tubercles and weakly curved rostrum. Also, penis and female sternite VIII are characteristic.