Cymosema Heraty & Burks

Burks, Roger A., Heraty, John M., Mottern, Jason, Dominguez, Chrysalyn & Heacox, Scott, 2017, Biting the bullet: revisionary notes on the Oraseminae of the Old World (Hymenoptera, Chalcidoidea, Eucharitidae), Journal of Hymenoptera Research 55, pp. 139-188: 150-151

publication ID

publication LSID

persistent identifier

taxon LSID

treatment provided by

Journal of Hymenoptera Research by Pensoft

scientific name

Cymosema Heraty & Burks

gen. n.

Cymosema Heraty & Burks   gen. n. Figs 15-20 View Figures 15–20 , 21-26 View Figures 21–26 , 27 View Figure 27

Type species.

Cymosema waterworthae   Burks & Mottern sp. n.


Based on the Greek κύμα (kyma) for wave and referring to the sinuate transscutal articulation; gender feminine.


The two species in this group are each known from single specimens collected in Australia, both among the smallest known Eucharitidae   . They differ from previously described species of Orasema   by having a cylindrical ventrally fused petiole that tapers basally to the articulating condyle (Figs 20 View Figures 15–20 , 25 View Figures 21–26 ), reduced mouthparts (Figs 16 View Figures 15–20 , 21 View Figures 21–26 ), and advanced axillae (Figs 19 View Figures 15–20 , 24 View Figures 21–26 ). The mesoscutum and mesoscutellum are separated by a distinct transscutal articulation, but the sclerites are tightly associated and the axillae are advanced (thus somewhat resembling the state in Timioderus   and Indosema   ). Based on morphological (reduced anellus, reduced mouthparts, smooth head and mesosoma) and molecular evidence, they are the potential sister group of Indosema   . The fore wing shape is unique within Eucharitidae   , with the anterior leading edge of the fore wing anteriorly curved at the stigmal vein giving the wing an oval-shaped appearance (Fig. 27 View Figure 27 ); in other Eucharitidae   the wing margin beyond the stigmal vein is straight (Fig. 7 View Figures 7–12 ). Male unknown.


Recognized from most Oraseminae  by the sinuate closely associated transscutal articulation, minute discoidal anellus, reduced mouthparts that are at most bidentate and medially directed (not falcate), basally tapered petiole and elongate slightly curved ovipositor. Distinguished from other Old World genera by the head transverse in frontal view and mostly smooth (Figs 16 View Figures 15–20 , 21 View Figures 21–26 ); scrobal depression strongly impressed and forming weak parallel channels, but without dorsal foveae; dorsal occipital margin abrupt and rounded; funicle 7-segmented; mandibles, chisel-shaped and slightly broader than long or bidentate with both teeth projecting medially (not falcate as in other Eucharitidae   ); labrum appears to be membranous and without digits; palpi greatly reduced; mesonotum appearing bare, at most with minute setae; mesoscutal lateral lobes entirely smooth or weakly reticulate laterally; frenum smooth; transscutal articulation sinuate and distinct but sclerites closely associated and appearing fused; callus bare; propodeal disc smooth (Fig. 20 View Figures 15–20 ) or weakly reticulate with single median carina (Fig. 24 View Figures 21–26 ); prepectus foveate and loosely articulated with pronotum ventrally (Fig. 18 View Figures 15–20 ); fore wing with basal area and specular area evenly covered with minute setae, and anterior margin of fore wing disc curved at stigmal vein (Fig. 27 View Figure 27 ); postmarginal vein short and about 1.5 × as long as stigmal vein; petiole base tapered and without transverse flange (Figs 20 View Figures 15–20 , 25 View Figures 21–26 ); petiole fused ventrally, basally tapering to condyle and without basal flange; acrosternite swollen and finely reticulate, antecostal sulcus not apparent, but a distinct transition to the smooth posterior region of Gs1; ovipositor long and slightly curved anteriorly (Fig. 26 View Figures 21–26 ); first valvula with 3-4 lateral teeth and with a strong oblique subapical ridge.

Host association.



Two species. Australia (Queensland and Western Australia).