Loncovilius variabilis, Reyes-Hernández & Hansen & Shaw & Solodovnikov, 2024

Reyes-Hernández, José L., Hansen, Aslak Kappel, Shaw, Josh Jenkins & Solodovnikov, Alexey, 2024, Phylogeny-based taxonomic revision and niche modelling of the rove beetle genus Loncovilius Germain, 1903 (Coleoptera: Staphylinidae: Staphylininae), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 202 (1), pp. 1-42 : 28

variabilis group

Included species: Loncovilius variabilis sp. nov..

Diagnosis: Head usually elongate, without coarse non-setiferous punctures; nuchal ridge complete dorsally, joined with infraorbital ridge; gula with isodiametric microsculpture in the middle; postmandibular ridge located close to eye margin ( Fig. 8F ); PMP (usually reduced); females with a11 distinctly longer than a10 (a11:a10 ratio>1.5). Pronotum slightly wider than long, its outline evenly curved. Elytra with even but sparse setiferous punctation on disc and epipleura. Protibiae not sexually dimorphic, in both sexes without laterodorsal row of thick spines. Mesotarsomeres 1–4 not sexually dimorphic, in both sexes with pale adhesive setae and mesotarsomeres 1–3 in both sexes with terminal plate. Tergites III and IV with several rows of fine punctures of moderate density, without sizeable impunctate area near to posterior margin ( Fig. 9B ); tergite VIII only slightly emarginate medio-apically in females; ovipositor, its second gonocoxite with one macroseta medially. Aedeagus as in Figure 10I–K .

Figures

Figure 8. A, pronotum of L. carlsbergi. AMRSP = anterior marginal row of setiferous punctures, APP = additional paired punctures adjacent to AMRSP, LLSP = large lateral setiferous puncture, LPSP = large posterior setiferous puncture, PPDS = paired punctures on dorsal serial, SLSP = sublateral setiferous puncture.B, L. cantharoides ventral view of mesotarsomere 3 with pale adhesive setae with terminal plate.C, L. cantharoides ventral view of metatarsomere 4, apical margin emarginated (bilobed), with pale adhesive setae without terminal plate. D, L. cantharoides tergites VI and VII.Black arrow up = anterior transverse basal carina (ATBC) laterally straight continuing to paratergites, white arrow down = ATBC descending laterally without reaching tergite margin.Head in ventral view of Loncovilius species. E, L. hammondi. F, L. lividipennis. PMP = postmandibular puncture, PMR = postmandibular ridge, EM = eye margin.

Figure 9. Details of the abdominal structures of Loncovilius species; tergites III and IV (A–E); ovipositor (F–H). A, Loncovilius germaini. B, L. variabilis. C, L. edwardsianus. D, L. semiflavus. E, L. lividipennis. F, L. impunctus. G, L. semiflavus. H, L. edwardsianus.

Figure 10. Aedeagus of Loncovilius species. A–C, Loncovilius germaini. D–F, L. hammondi. G, H, L. impunctus. I–K, L. variabilis. L–N, L. edwardsianus. O–Q, L. cantharoides. A, D, G, I, L, O, aedeagus dorsally (parameral side). B, E, J, M, P, apical portion of paramere, underside. C, F, H, K, N, Q, aedeagus laterally.