Laricobius laticollis Fall, 1916

Leschen, Richard A. B., 2011, World review of Laricobius (Coleoptera: Derodontidae), Zootaxa 2908, pp. 1-44 : 15-16

10. Laricobius laticollis Fall, 1916

(figs 11, 41)

Diagnosis. Body bicolored. Head dark; ocelli present. Prothorax transverse with well developed lateral carina and sides explanate. Pronotum light; posterior tooth present or absent. Scutellum usually dark. Surface of elytra weakly incurvate; elytral punctures not confluent. Epipleura dark. Ventrites dark. Femora light. Tibiae light. Aedeagus without median carina on phallobase; medium lobe apiculate; apices of parameres without an internal ridge.

Description. Length 2.30–2.76 mm (x = 2.52, n = 15). Body bicolored with head, scutellum (usually), epipleuron, and venter chocolate brown; pronotum, elytra, and hypomeron light brown or tan; antenna light brown or tan with AI usually darker, legs reddish light brown or tan, bases and apices of the femora and tibiae may be darker. Head with ocelli; u-shaped furrow present; interocular distance about 4x the width of the eye; macropunctures rounded and distinct, sometimes fusing and sometimes present at the center of the frons; micropunctures coarse; setae erect, moderately elongate with an average length as long as 2/3 the width of the eye. Antenna with length of A3 about equal to A4, ratios 2.1:1.6:1.3:1.4:1.4:1.1:1.1:1.0:1.6:1.4:2.1; A11 not strongly asymmetrical. Prothorax transverse and rectangular, dorsoventrally flattened, and widest at middle, sides converging anteriorly from widest point and weakly convex posteriorly; about 0.68 x as long as wide (pronotal length/greatest pronotal width = 0.63– 0.71, x = 0.68); depth = 0.26–0.40 mm (x = 0.32); pronotum laterally explanate, lateral carina with a distinct sharp edge; anterior angle about 60° and subacute, not forming a tooth; foveae visible in dorsal view; posterior angle indistinct or distinct with short tooth present or absent; macropunctures on disc distinct, well-impressed and wellseparated or contiguous at the anterior middle of disc, micropunctures coarse; setae suberect, short and average length about 1/2 width of eye. Elytra about 3.22 x as long as wide (elytral length/greatest elytral width = 3.00–3.47, x = 3.22) and 3.35 x as long as pronotum (elytral length/pronotal length = 3.20–3.54, x = 3.35); surface weakly incurvate at basal third; macropunctures not coalescing to form grooves (apart from those of striae 1 along the suture near apex in some specimens), separated by an average of 1/2 puncture diameter; micropunctures fine; microsculpture visible on some specimens; setae suberect, short and an average of about 1/2 the length of the eye. Aedeagus with phallobase transverse, about 1/2 the length of the median lobe, median carina absent; median lobe projecting slightly beyond apices of parameres, moderately broad with its greatest width subequal to parameres, sides gradually narrowing posteriorly with the apex apiculate, median groove or carina absent; ostium subapical; parameres broad and rounded apically, short setae present on apices and along inner margin, internal subapical ridge absent. Spiculum gastrale apically narrowed.

Comments. Among the species having an explanate pronotal margin, L. laticollis is one of three species with the head dark and pronotum light. It can be distinguished from L. caucasicus by having an incurvate elytral surface and from L. baoxingensis by the shape of the pronotum.

Fall (1916) had 14 specimens, taken at the University of Washington Campus February and April, but I have only examined specimens collected in February 1911. The single specimen labelled as “ type ” is here designated as the lectotype, this does not have a black scutellum; the remaining syntypes are mixed.

Hosts. Plants: Pseudotsuga menzeseii (Mirb.) Franco , Tsuga heterophylla (Raf.) Sarg. Adelgid : Adelges cooleyi (Gillette) (M. Montgomery, pers. com.).

Distribution. United States: California, Idaho, Oregon, Washington. Canada: Alberta, British Columbia.

Type material examined. 7 syntypes (all pointed MCZ): 1, Seattle, II-12-11 (hw)/H. C. FALL COLLEC- TION /W. T./ TYPE laticollis . (name hw)/ M.C.Z. Type 24993 (red label, number hw)/ Aug–Dec 2005 MCZ Image Database (bordered label with camera icon)/ LECTOTYPE, designated by R. Leschen 2009; 1, W. T./Univ. of W. Campus. Seattle. IV–6–12 (hw)/ H. C. FALL COLLECTION/ Laricobius laticollis Fall. (hw); 2, Seattle Wash. (hw)/ H. C. FALL COLLECTION; 1, W. T./Seattle, Campus Univ. of W., II-12-11 (hw)/ H. C. FALL COLLEC- TION; 1, W. T./Seattle, II-12-11 (hw)/ H. C. FALL COLLECTION; 1, “Campus” (hw), Seattle, Wash (printed) II- 12-11 T. C. K. (hw)/ H. C. FALL COLLECTION.

Additional material examined. CANADA. Alberta: 1, Truner V., Alta. VI.1.1933, Owen Bryant ( CAS); 20, Ghost Dam, Alberta, several dates, BF and JL Carr ( CNC); 2, Calgary, 1973, BF and JL Carr ( CNC); 1, Coleman, Alberta ( CNC). British Columbia: 2, Salmon Arm, 3.V.35, Hugh Leech, 10351; 1, same but 30.IV.1930 ( CAS); 1, same but 30.IV.1933 on Douglas fir boughs, 23.III.34, beating Douglas fir branches, E. S. Ross Collection ( CAS); 1, Sanca, 22.IV.1933 G. Stace Smith, from swift stream (1, CAS; 4, CNC). UNITED STATES. California: 8, Fieldbrook, Humboldt Co., Mar.27, ’38, B. P. Bliven No. 801( CAS); Humbolt Co., v.3.17, F.W. Nunenmacher ( FMNH). Idaho: 1, Moscow, v-28 1950, N.M. Downie ( FMNH); 1, same but v-20, 1950 ( FMNH); 1, Troy, v-14, 1950, N.M. Downie ( FMNH); 1, Deary, v-14, 1950, N.M. Downie ( FMNH); 1, Moscow Mts, II.10.1934, I.W. Bales ( FMNH). Oregon: 7, Milton, Ore, G. H. Nelson, IV-3-49, G. W. Green collection ( CAS). Washington: 1, Fort Lewis, Pierce Co. V-3.1946, P. H. arnaudf ( CAS); 2, Seattle, VII-13 on Douglas Fir, Van Dyke Collection ( CAS); 1, Seattle, Wash. III-7-13, sweeping Douglas fir ( CAS); 3, Grand Coulee, Washington, BF and JL Carr ( CNC); 11, King Co., Seattle; 17-24 February 2007, D. McDonald, Havill Samples # 07-01, 07-01A, 07-01B, 07-01.1 ( YPM).