Escaryus retusidens Attems, 1904

Dyachkov, Yurii V. & Bonato, Lucio, 2024, An updated synthesis of the Geophilomorpha (Chilopoda) of Asian Russia, ZooKeys 1198, pp. 17-54 : 17

35. Escaryus retusidens Attems, 1904

Escaryus retusidens Attems 1904: 121.

Escaryus retusidens - Attems 1929: 96; Titova 1972: 135, 116; 1973: 110; Volkova 2016: 675; Zuev 2016: 33; Nefediev et al. 2017a: 11; 2017c: 222; 2018: 239; Dyachkov and Tuf 2018: 295; Nefediev 2019: 27; Dyachkov 2022: 25.

Type locality.

Kyrgyzstan: Issyk-Kul Region: “Przewalsk” ( Attems 1904) = Karakol.

Type series.

Syntypes: 4 specimens, including 2 males and 2 females. Deposited in NHMW ( Ilie et al. 2009).

Diagnosis.

An Escaryus species with body length reaching ≥ 4 cm; clypeus with small plagulae; labral arc relatively deep, with denticles long and obtuse; first maxillae with one pair of lappets; forcipular trochanteroprefemur, femur, and tibia with small denticles, tarsungulum with a small bulge; 49-55 leg-bearing segments; metasternites with relatively sparse setae; metasternite of the ultimate leg-bearing segment trapezoid, approximately as long as wide; coxal pores on both ventral and lateral sides of coxopleura; anal pores absent.

Distribution.

Western Siberia: Altai krai, Republic of Altai, and Kemerovo oblast ( Nefediev et al. 2017a, 2017c, 2018; Nefediev 2019). Possibly Far East ( Titova 1972, 1973; see Remarks). Outside Asian Russia: westwards to Moldova ( Titova 1973; Volkova 2016; Zuev 2016; Zuev and Evsyukov 2016); southwards to Kazakhstan and Kyrgyzstan ( Attems 1904; Titova 1972, 1973; Dyachkov and Tuf 2018; Dyachkov 2022).

Remarks.

A total of 19 males and 26 females collected from Trans-Ili Alatau (Almaty Region of Kazakhstan) were indicated by Titova (1973: 110) as lectotypes, but this action is not valid, as the syntypes still exist ( Ilie et al. 2009).

Titova (1972: 135; 1973: 116) wrote "... it is possible to distinguish territories from Cisamuria to Kuzbass and Altai, where 4 species are spread: E. koreanus , E. japonicus , E. retusidens , and E. chadaevae ". However, she did not mention material from this area when she listed the studied material of E. retusidens ( Titova 1973: 110): "Kazakhstan, Trans-Ili Alatau … Moreover, E. retusidens were studied from the Dzhungarian Alatau, the Greater Caucasus, Crimea, Moldova, Rostov, Voronezh and Voroshilovograd oblasts".