Triozidae

Burckhardt, Daniel & Ouvrard, David, 2012, A revised classification of the jumping plant-lice (Hemiptera: Psylloidea), Zootaxa 3509, pp. 1-34: 20-23

publication ID

http://doi.org/ 10.5281/zenodo.209825

publication LSID

lsid:zoobank.org:pub:16A98DD0-D7C4-4302-A3B7-64C57768A

persistent identifier

http://treatment.plazi.org/id/C46C5928-776B-FFE2-0385-9220FDEAFB47

treatment provided by

Plazi

scientific name

Triozidae
status

 

Triozidae  

Triozinae Löw, 1879: 605, 609.

Pauropsyllinae Crawford, 1914: 42.

Siphonaleyrodinae Takahashi, 1932: 48.

Bactericerini Heslop-Harrison, 1958: 577, 578.

Eutriozini Loginova, 1964: 473.

Hemischizocraniini Bekker-Migdisova, 1973: 115.

Leptinopterinae (sic) Bekker-Migdisova, 1973: 104.

Paracomecini Bekker-Migdisova, 1973: 115.

Epitriozini Kwon, 1984: 79.

Trichochermini Kwon, 1983: 82.

Neolithinae White & Hodkinson, 1985: 273.

Rhinopsyllidae Klimaszewski, 1993: 65.

Carsitriidae Li, 2011: 1303; syn. nov.

Neotriozidae Li, 2011: 1307; syn. nov.

Triozopsinae Li, 2011: 1383; syn. nov.

Asiotriozinae Li, 2011: 1512; syn. nov.

Metatriozidinae Li, 2011: 1513; syn. nov. Diagnosis: Hollis (1984), Burckhardt (1988), Burckhardt and Lauterer (1997 b).

Internal relationships: White and Hodkinson (1985) recognised the three subfamilies Neolithinae,

Triozamiinae and Triozinae. Hollis and Broomfield (1989) transferred the Triozamiinae to the Homotomidae   . The

erection of the Neolithinae by White and Hodkinson (1985) is based on a misidentification of a larva which is

referrable to a species related to Neotrioza tavaresi Crawford   rather than Neolithus   (D. Burckhardt, unpubl.).

Schedoneolithus   , the only other member of the subfamily shares the large frons but shows otherwise no detailed

synapomorphies with Neolithus   , making the monophyly of Neolithinae doubtful. White and Hodkinson (1985)

defined the Triozinae by the presence of genal processes, the anal break being in distance from the apex of vein

Cu 1 b in the forewing and the lack of a rhinarium on antennal segment 3 in the adults as well as the dorsally

completely fused sclerites of head and thorax in the larva. The first character is variable in many triozid genera and

the other three are present also in members of the Neolithinae, leaving the subfamily undefined and, therefore,

paraphyletic. Recently Li (2011) proposed a classification treating the triozids as a superfamily with four families

and several subfamilies. His artificial classification based on the Chinese fauna does not reflect phylogeny. In

conclusion, there is currently no scientifically convincing suprageneric classification for Triozidae   . Comments: This is a species-rich, probably monophyletic family. In contrast, most of the genera are ill-

defined and artificial, and the phylogenetic relationships between genera remain largely unknown. Here we do not

use further subdivisions into subfamilies and tribes as none of the currently proposed groupings is supported by

convincing characters. The use of the generic names varies greatly between authors. Here we list many genera as

valid which are considered synonyms by some authors. This is particularly the case with genera synonymised with

Trioza   which is treated, by most authors, as an artificial receptacle for species not showing any particular

morphological modifications. A sound phylogenetic analysis is required for obtaining a better base for a stable

generic classification.

Aacanthocnema Tuthill & Taylor, 1955: 252   ; type-species: Trioza casuarinae Froggatt, 1901   , by original designation.

Acanthocasuarina Taylor   in Taylor et al. (2011): 27; type-species: Acanthocasuarina acutivalvis Taylor   in Taylor et al. (2011), by original designation.

Afrotrioza Hollis, 1984: 25   ; type-species: Afrotrioza bersama Hollis, 1984   , by original designation and monotypy.

Anomocephala Tuthill, 1942: 75   ; type-species: Anomocephala unica Tuthill, 1942   , by original designation and monotypy.

Asiotrioza Li, 2011: 1512, 1900; type-species: Asiotrioza zhongfeiensis Li, 2011, by original designation and monotypy.

Bactericera Puton, 1876: 286; type-species: Bactericera perrisii Puton, 1876, by monotypy (= Allotrioza, Carsitria syn. nov., Eubactericera, Klimaszewskiella replacement name for Smirnovia Klimaszewski, 1968, Paratrioza, Rhinopsylla)

Baeoalitriozus Li, 2011: 1356; type-species: Baeoalitriozus yangi Li, 2011, by original designation and monotypy.

Berchemitrioza Li, 2011: 1386, 1886; type-species: Berchemitrioza acutata Li, 2011, by original designation.

Calinda Blanchard, 1852: 309   ; type-species: Calinda testacea Blanchard, 1852   , by subsequent designation of Burckhardt (1988).

Casuarinicola Taylor   in Taylor et al. (2010): 3; type-species: Casuarinicola australis Taylor   in Taylor et al. (2010), by original designation.

Cecidotrioza Kieffer, 1908: 159   ; type-species: Cecidotrioza baccarum Kieffer, 1908   , by monotypy; (= Homotrioza   ).

Ceropsylla Riley, 1885: 76   : type-species: Ceropsylla sideroxyli Riley, 1884   , by monotypy.

Cerotrioza Crawford, 1918: 454   ; type-species: Cerotrioza bivittata Crawford, 1918   , by original designation and monotypy.

Chouitrioza Li, 1989: 217, 221 type-species: Chouitrioza sinica Li, 1989, by original designation and monotypy.

Colopelma Enderlein, 1926: 400; type-species: Trioza thomasii Löw, 1888, by original designation.

Conicotrioza Li, 2005: 193, 211; type-species: Conicotrioza cylindrata Li, 2005, by original designation and monotypy.

Crawforda Caldwell, 1940: 397; type-species: Crawforda triopsyllina Caldwell, 1940, by original designation and monotypy.

Dolichotrioza Li, 2002: 172, 187; type-species: Dolichotrioza sinuata Li, 2002, by original designation and monotypy.

Dyspersa Klimaszewski, 1968: 11   ; type-species: Trioza apicalis Foerster, 1848   , by original designation.

Egeirotrioza Boselli, 1931: 268   ; type-species: Trioza ceardi   de Bergevin, 1926, by original designation and monotypy; (= Evegeirotrioza syn. nov.)

Engytatoneura Loginova, 1972: 33   ; type-species: Engytatoneura lindbergi Loginova, 1972   , by original designation and monotypy.

Eotrioza Konovalova, 1987: 34   ; type-species: Eotrioza ussuriensis Konovalova, 1987   , by original designation and monotypy.

Epitrioza Kuwayama, 1910: 55   ; type-species: Epitrioza mizuhonica Kuwayama, 1910   , by original designation and monotypy.

Eryngiofaga Klimaszewski, 1968: 10   ; type-species: Trioza mesomela Flor, 1861   , by original designation.

Eutrioza Loginova, 1964: 474   ; type-species: Eutrioza opima Loginova, 1964   , by monotypy.

Furcitrioza Li, 2011: 1348, 1880; type-species: Furcitrioza lecanthae Li, 2011, by original designation and monotypy.

Genotriozus Li, 2011: 1366, 1884; type-species: Genotriozus octoimaculatus Li, 2011, by original designation.

Hemischizocranium Tuthill, 1956: 158   ; type-species: Hemischizocranium bessi Tuthill, 1956   , by original designation and monotypy.

Hemitrioza Crawford, 1914: 104   ; type-species: Hemitrioza sonchi Crawford, 1914   , by original designation and monotypy.

Heterotrioza Dobreanu & Manolache, 1960: 94   ; type-species: Trioza obliqua Thomson, 1877   , by monotypy (= Trioza (Halotrioza)   syn. nov., Triozidus   ).

Hevaheva Kirkaldy, 1902: 113   ; type-species: Hevaheva perkinsi Kirkaldy, 1902   , by monotypy.

Hippophaetrioza Conci & Tamanini, 1984: 240   ; type-species: Trioza binotata Löw, 1883   , by original designation (= Hippophaetrioza (Maculatrioza)   syn. nov.).

Izpania Klimaszewski, 1962: 254   ; type-species: Izpania acona Klimaszewski, 1962   , by original designation and monotypy.

Kuwayama Crawford, 1911: 503   ; type-species: Paratrioza medicaginis Crawford, 1910   , by original designation; (= replacement name for Epitrioza, Succinopsylla   † syn. nov.).

Lauritrioza Conci & Tamanini, 1986: 238   ; type-species: Trioza alacris Flor, 1861   , by original designation and monotypy.

Leptotrioza Miyatake, 1972: 29   ; type-species: Neotriozella bicolor Crawford, 1915   , by original designation and monotypy.

Leptynoptera Crawford, 1919: 147   ; type-species: Leptynoptera sulfurea Crawford, 1919   , by original designation and monotypy.

Leuronota Crawford, 1914: 67   ; type-species: Trioza maculata Crawford, 1910   , by original designation; (= Paracomeca   ).

Levidea Tuthill, 1938: 245   ; type-species: Levidea lineata Tuthill, 1938   , by original designation and monotypy.

Megatrioza Crawford, 1915: 264   ; type-species: Megatrioza armata Crawford, 1915   , by original designation and monotypy.

Metatrioza Tuthill, 1939: 53   ; type-species: Metatrioza pubescens Tuthill, 1939   , by original designation and monotypy.

Neolithus Scott, 1882: 445   ; type-species: Neolithus fasciatus Scott, 1882   , by monotypy.

Neotrioza Kieffer, 1905: 175   ; type-species: Neotrioza machili Kieffer, 1905   , by monotypy.

Neotriozella Crawford, 1911: 503   ; type-species: Trioza immaculata Crawford, 1910   , by original designation; (= replacement name for Neotrioza   ).

Ozotrioza Kieffer, 1905: 178   ; type-species: not designated.

Parastenopsylla Yang, 1984: 202   ; type-species: Stenopsylla proboscidaria Yu, 1956   , by original designation; (= Indotrioza   ).

Pariaconus Enderlein, 1926: 401   ; type-species: Kuwayama nigricapita Crawford, 1918   , by original designation.

Pauropsylla Rübsaamen, 1899: 264   ; type-species: Pauropsylla udei Rübsaamen, 1899   , by monotypy; (= Neotrioza sensu Li, 2011: 1313   , nec Kieffer, 1905: 175, misinterpretation, Sympauropsylla   ).

Paurotriozana Caldwell, 1940: 396   ; type-species: Paurotriozana adaptata Caldwell, 1940   , by original designation and monotypy.

Petalolyma Scott, 1882: 459   ; type-species: Psylla basalis Walker, 1858   , by monotypy.

Phylloplecta Riley, 1884: 319   ; type-species: Psylla tripunctata Fitch, 1851   , by monotypy; (= Choricymoza   , Sinitrioza   ).

Powellia Maskell, 1879: 223   ; type-species: Powellia vitreoradiata Maskell, 1879   , by monotypy.

Pseudotrioza Miyatake, 1972: 27   ; type-species: Pseudotrioza hiurai Miyatake, 1972   , by original designation and monotypy.

Rhegmoza Enderlein, 1918: 480   ; type-species: Rhegmoza tinctoria Enderlein, 1918   , by original designation and monotypy.

Rhinopsyllida   † Klimaszewski, 1997: 37; type-species: Rhinopsyllida acutealla   † Klimaszewski, 1997, by original designation and monotypy.

Schedoneolithus Tuthill, 1959: 26   ; type-species: Schedoneolithus dunaliae Tuthill, 1959   , by original designation and monotypy.

Schedotrioza Tuthill & Taylor, 1955: 251   ; type-species: Trioza eucalypti Froggatt, 1901   , by original designation.

Siphonaleyrodes Takahashi, 1932: 48   ; type-species: Siphonaleyrodes formosanus Takahashi, 1932   , by original designation and monotypy.

Spanioza Enderlein, 1926: 400   ; type-species: Trioza galii Foerster, 1848   , by original designation.

Stenopsylla Kuwayama, 1910: 53   ; type-species: Stenopsylla nigricornis Kuwayama, 1910   , by original designation and monotypy (= Cryptotrioza   , Dasymastix   , Eustenopsylla, Philippinocarsia   syn. nov.).

Swezeyana Caldwell, 1940: 389   ; type-species: Swezeyana elongagena Caldwell, 1940   , by original designation.

Torulus Li, 1991: 37, 41; type-species: Torulus sinicus Li, 1991, by original designation and monotypy.

Trachotrioza Li, 2011: 1350, 1881; type-species: Trachotrioza beijingensis Li, 2011, by original designation.

Trichochermes Kirkaldy, 1904: 280   ; type-species: Trioza walkeri Foerster, 1848   , by original designation; (= replacement name for Trichopsylla   ).

Trioacantha   † Klimaszewski, 1998: 23; type-species: Trioacantha indocilia   † Klimaszewski, 1998, by original designation and monotypy.

Trioza Foerster, 1848: 67   , 82; type-species: Chermes urticae Linné, 1758   , by subsequent designation of Oshanin (1912); (= Metatriozidus   , Triozopsis   ).

Trisetitrioza Li, 1995: 21, 25; type-species: Trisetitrioza clavellata Li, 1995, by original designation and monotypy; (= Neorhinopsylla).

Triozoida Crawford, 1911: 491   ; type-species: Triozoida johnsonii Crawford, 1911   , by original designation; (= Myrmecephala   , Optomopsylla   ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Triozidae

Loc

Triozidae

Burckhardt, Daniel & Ouvrard, David 2012
2012
Loc

Neotrioza sensu

Li 2011: 1313
2011
Loc

Trioacantha

Klimaszewski 1998: 23
1998
Loc

Trioacantha indocilia

Klimaszewski 1998
1998
Loc

Rhinopsyllida

Klimaszewski 1997: 37
1997
Loc

Rhinopsyllida acutealla

Klimaszewski 1997
1997
Loc

Eotrioza

Konovalova 1987: 34
1987
Loc

Eotrioza ussuriensis

Konovalova 1987
1987
Loc

Lauritrioza

Conci & Tamanini 1986: 238
1986
Loc

Afrotrioza

Hollis 1984: 25
1984
Loc

Afrotrioza bersama

Hollis 1984
1984
Loc

Hippophaetrioza

Conci & Tamanini 1984: 240
1984
Loc

Parastenopsylla

Yang 1984: 202
1984
Loc

Engytatoneura

Loginova 1972: 33
1972
Loc

Engytatoneura lindbergi

Loginova 1972
1972
Loc

Leptotrioza

Miyatake 1972: 29
1972
Loc

Pseudotrioza

Miyatake 1972: 27
1972
Loc

Pseudotrioza hiurai

Miyatake 1972
1972
Loc

Dyspersa

Klimaszewski 1968: 11
1968
Loc

Eryngiofaga

Klimaszewski 1968: 10
1968
Loc

Eutrioza

Loginova 1964: 474
1964
Loc

Eutrioza opima

Loginova 1964
1964
Loc

Izpania

Klimaszewski 1962: 254
1962
Loc

Izpania acona

Klimaszewski 1962
1962
Loc

Heterotrioza

Dobreanu & Manolache 1960: 94
1960
Loc

Schedoneolithus

Tuthill 1959: 26
1959
Loc

Schedoneolithus dunaliae

Tuthill 1959
1959
Loc

Hemischizocranium

Tuthill 1956: 158
1956
Loc

Hemischizocranium bessi

Tuthill 1956
1956
Loc

Stenopsylla proboscidaria

Yu 1956
1956
Loc

Aacanthocnema

Tuthill & Taylor 1955: 252
1955
Loc

Schedotrioza

Tuthill & Taylor 1955: 251
1955
Loc

Anomocephala

Tuthill 1942: 75
1942
Loc

Anomocephala unica

Tuthill 1942
1942
Loc

Paurotriozana

Caldwell 1940: 396
1940
Loc

Paurotriozana adaptata

Caldwell 1940
1940
Loc

Swezeyana

Caldwell 1940: 389
1940
Loc

Swezeyana elongagena

Caldwell 1940
1940
Loc

Metatrioza

Tuthill 1939: 53
1939
Loc

Metatrioza pubescens

Tuthill 1939
1939
Loc

Levidea

Tuthill 1938: 245
1938
Loc

Levidea lineata

Tuthill 1938
1938
Loc

Siphonaleyrodes

Takahashi 1932: 48
1932
Loc

Siphonaleyrodes formosanus

Takahashi 1932
1932
Loc

Egeirotrioza

Boselli 1931: 268
1931
Loc

Pariaconus

Enderlein 1926: 401
1926
Loc

Spanioza

Enderlein 1926: 400
1926
Loc

Leptynoptera

Crawford 1919: 147
1919
Loc

Leptynoptera sulfurea

Crawford 1919
1919
Loc

Cerotrioza

Crawford 1918: 454
1918
Loc

Cerotrioza bivittata

Crawford 1918
1918
Loc

Kuwayama nigricapita

Crawford 1918
1918
Loc

Rhegmoza

Enderlein 1918: 480
1918
Loc

Rhegmoza tinctoria

Enderlein 1918
1918
Loc

Neotriozella bicolor

Crawford 1915
1915
Loc

Megatrioza

Crawford 1915: 264
1915
Loc

Megatrioza armata

Crawford 1915
1915
Loc

Hemitrioza

Crawford 1914: 104
1914
Loc

Hemitrioza sonchi

Crawford 1914
1914
Loc

Leuronota

Crawford 1914: 67
1914
Loc

Kuwayama

Crawford 1911: 503
1911
Loc

Neotriozella

Crawford 1911: 503
1911
Loc

Triozoida

Crawford 1911: 491
1911
Loc

Triozoida johnsonii

Crawford 1911
1911
Loc

Epitrioza

Kuwayama 1910: 55
1910
Loc

Epitrioza mizuhonica

Kuwayama 1910
1910
Loc

Paratrioza medicaginis

Crawford 1910
1910
Loc

Trioza maculata

Crawford 1910
1910
Loc

Trioza immaculata

Crawford 1910
1910
Loc

Stenopsylla

Kuwayama 1910: 53
1910
Loc

Stenopsylla nigricornis

Kuwayama 1910
1910
Loc

Cecidotrioza

Kieffer 1908: 159
1908
Loc

Cecidotrioza baccarum

Kieffer 1908
1908
Loc

Neotrioza

Kieffer 1905: 175
1905
Loc

Neotrioza machili

Kieffer 1905
1905
Loc

Ozotrioza

Kieffer 1905: 178
1905
Loc

Trichochermes

Kirkaldy 1904: 280
1904
Loc

Hevaheva

Kirkaldy 1902: 113
1902
Loc

Hevaheva perkinsi

Kirkaldy 1902
1902
Loc

Trioza casuarinae

Froggatt 1901
1901
Loc

Trioza eucalypti

Froggatt 1901
1901
Loc

Pauropsylla Rübsaamen, 1899: 264

Rubsaamen 1899: 264
1899
Loc

Pauropsylla udei Rübsaamen, 1899

Rubsaamen 1899
1899
Loc

Ceropsylla

Riley 1885: 76
1885
Loc

Ceropsylla sideroxyli

Riley 1884
1884
Loc

Phylloplecta

Riley 1884: 319
1884
Loc

Trioza binotata Löw, 1883

Low 1883
1883
Loc

Neolithus

Scott 1882: 445
1882
Loc

Neolithus fasciatus

Scott 1882
1882
Loc

Petalolyma

Scott 1882: 459
1882
Loc

Powellia

Maskell 1879: 223
1879
Loc

Powellia vitreoradiata

Maskell 1879
1879
Loc

Trioza obliqua

Thomson 1877
1877
Loc

Trioza mesomela

Flor 1861
1861
Loc

Trioza alacris

Flor 1861
1861
Loc

Psylla basalis

Walker 1858
1858
Loc

Calinda

Blanchard 1852: 309
1852
Loc

Calinda testacea

Blanchard 1852
1852
Loc

Psylla tripunctata

Fitch 1851
1851
Loc

Trioza apicalis

Foerster 1848
1848
Loc

Trioza galii

Foerster 1848
1848
Loc

Trioza walkeri

Foerster 1848
1848
Loc

Trioza

Foerster 1848: 67
1848
Loc

Chermes urticae Linné, 1758

Linne 1758
1758