Hultholia Gagnon & G. P. Lewis, PhytoKeys 71: 58. 2016.

Bruneau, Anne, de Queiroz, Luciano Paganucci, Ringelberg, Jens J., Borges, Leonardo M., Bortoluzzi, Roseli Lopes da Costa, Brown, Gillian K., Cardoso, Domingos B. O. S., Clark, Ruth P., Conceicao, Adilva de Souza, Cota, Matheus Martins Teixeira, Demeulenaere, Else, de Stefano, Rodrigo Duno, Ebinger, John E., Ferm, Julia, Fonseca-Cortes, Andres, Gagnon, Edeline, Grether, Rosaura, Guerra, Ethiene, Haston, Elspeth, Herendeen, Patrick S., Hernandez, Hector M., Hopkins, Helen C. F., Huamantupa-Chuquimaco, Isau, Hughes, Colin E., Ickert-Bond, Stefanie M., Iganci, Joao, Koenen, Erik J. M., Lewis, Gwilym P., de Lima, Haroldo Cavalcante, de Lima, Alexandre Gibau, Luckow, Melissa, Marazzi, Brigitte, Maslin, Bruce R., Morales, Matias, Morim, Marli Pires, Murphy, Daniel J., O'Donnell, Shawn A., Oliveira, Filipe Gomes, Oliveira, Ana Carla da Silva, Rando, Juliana Gastaldello, Ribeiro, Petala Gomes, Ribeiro, Carolina Lima, Santos, Felipe da Silva, Seigler, David S., da Silva, Guilherme Sousa, Simon, Marcelo F., Soares, Marcos Vinicius Batista & Terra, Vanessa, 2024, Advances in Legume Systematics 14. Classification of Caesalpinioideae. Part 2: Higher-level classification, PhytoKeys 240, pp. 1-552 : 1

publication ID	https://dx.doi.org/10.3897/phytokeys.240.101716
persistent identifier	https://treatment.plazi.org/id/C3D26874-7A3F-3439-CBB7-B434BADBBD0E
treatment provided by	PhytoKeys by Pensoft
scientific name	Hultholia Gagnon & G. P. Lewis, PhytoKeys 71: 58. 2016.
status

Treatment

Hultholia Gagnon & G. P. Lewis, PhytoKeys 71: 58. 2016. View in CoL

Figs 35 View Figure 35 , 37 View Figure 37 , 60 View Figure 60

Type.

Hultholia mimosoides (Lam.) Gagnon & G. P. Lewis [≡ Caesalpinia mimosoides Lam.]

Description.

Climbing woody shrub; branches densely armed with short, robust, needle-like trichomes; young stems pubescent, with rust-coloured, hyaline hairs, and dome-shaped glands, topped with a few hairs. Stipules subulate, caducous. Leaves bipinnate; pinnae in 10-30 opposite pairs, with a pair of deflexed prickles at the insertion of the pinnae on the leaf rachis and at the insertion of leaflets on the pinnae rachides; leaflets 7-20 pairs per pinna, opposite, glabrous, eglandular. Inflorescence a terminal or leaf-opposed, lax raceme, with 50 or more flowers. Flowers bisexual, zygomorphic; hypanthium persisting as a wide shallow cup at the pedicel apex as fruit matures; sepals 5, caducous, the lower sepal strongly cucullate; petals 5, free, bright yellow, dark glands present on the blade, median petal smaller than the 4 lateral petals; stamens 10, free, filaments pubescent at least on the lower half; ovary densely pubescent and with glandular dots (often obscured by the dense pubescence). Fruit an obovoid, falcate, vesicular, unarmed, dehiscent legume, sparsely pubescent, particularly along the margin, and with a few obscure stellate hairs, and covered in gland dots, 1-3-seeded. Seeds sub-globose, grey.

Chromosome number.

2 n = 24 ( Fedorov 1969).

Included species and geographic distribution.

Monospecific ( H. mimosoides ), distributed across Asia, in China (Yunnan), Bangladesh, India, Laos, Myanmar (Burma), Thailand and Vietnam (Fig. 60 View Figure 60 ).

Ecology.

In secondary thickets and clearings, often on roadsides.

Etymology.

The name Hultholia honours the Cambodian botanist Dr. Sovanmoly Hul Thol (born 1946), whose doctoral thesis, "Contribution à la révision de quelques genres de Caesalpiniaceae , representés en Asie" (Hul Thol 1976), is an important revision of the Asian species and genera of the Caesalpinieae , and particularly the genus Pterolobium .

Human uses.

Although Hultholia mimosoides is not known to be cultivated, the young, pungent, flowering shoots are sold as a vegetable in markets in Vientiane (Laos) (Vidal and Hul Thol 1976).

Notes.

This monospecific genus was described by Gagnon et al. (2016) and has an unresolved phylogenetic position within a clade of lianescent Old World genera.

Taxonomic references.

Chen et al. (2010a); Gagnon et al. (2016); Vidal and Hul Thol (1976).