Ancylomenes tosaensis ( Kubo, 1951 )

Bruce, A. J., 2010, Ancylomenes kuboi and A. okunoi spp. nov. (Decapoda: Pontoniinae), from the Australian Northwest Shelf, Vietnam and the Philippines *, Zootaxa 2372 (1), pp. 169-176 : 174-175

Ancylomenes tosaensis ( Kubo, 1951)

( Fig. 4 )

Periclimenes (Ancylocaris) tosaensis Kubo, 1951: 268–271 , figs. 7–8.

Periclimenes tosaensis . — Hayashi 1986: 100–101, 261, pl. 61. — Li 2000: 243–244, fig. 323. — Li et al. 2005: 548– 549, fig. 33. — Bruce 2008; 15–16, fig. 9, 14D.

Ancylomenes tosaensis . — Okuno & Bruce, 2010: 101 .

Material examined. (i) 1 male, CL 3.3 mm, FRV Soela, stn NWS–16, B4, Northwest Shelf , Western Australia, 19°04.3’S 118°50.5’E, 80 m, sledge, 27 April 1983, NTM Cr015053 .

Remarks. The single specimen has a rostral dentition 1+8/2 ( Fig. 4A ), with the epigastric tooth separated by a larger interval from the first rostral tooth than between remaining teeth, both distal ventral teeth very small, exactly as illustrated by Li et al. (2005): plumose ventral setae particularly long ( Fig. 4A ). The proximal dorsal surface of the stylocerite also has numerous plumose setae. The basicerite has a small acute disto-dorsal tooth ( Fig. 4A ). The interocular process is small and rounded. The single second pereiopod has the fingers subequal to the palm length, dactylus and fixed finger both unarmed; the carpus also subequal to the palm length. The third ambulatory pereiopod has the dactylus ( Fig. 4C ) about 0.22 of the propodus length, 5.4 times longer than the proximal width, dorsal margin smoothly convex, without setae, ventral margin smoothly concave, unguis not clearly demarcated, about 0.3 of corpus length; propodus ( Fig. 4B ) about 0.68 of CL, 13.5 times longer than deep, disto-ventral angle oblique, with several simple setae disto-dorsally and a pair of slender unequal pre-terminal spines proximo-ventrally, lateral spine almost half the dactylus length, twice distal propodus width, medial spine about 0.6 of medial spine length, ventral margin with distal pair of one long and one very short spines and two small single spines.

Colouration. (i) From field notes: mainly transparent, rostrum transparent, third abdominal tergite with anterior red spot and posterior white transverse bar, antennular peduncle with yellow spots distally on each segment, scaphocerite transparent, first pereiopod similar to second, purple much less intense, second pereiopod chelae with palm white, hinge region and proximal part of fingers deep purple, more distal fingers feebly purplish, carpus yellow, distally purple, merus and ischium similar, ambulatory pereiopods with dactyli purple, propodi yellowish outlined with purple-red, coxae pale yellow, distal two thirds of telson white, with anterolateral red patch; uropodal exopod with distal two thirds white, with large distal purple eyespot, endopod with distal three fourths white, proximal fourth reddish.

Distribution. Type locality: Usa, Tosa Bay, Shikoku, Japan. Previously reported from the Northwest Shelf, from 100 m by Bruce (2008). Also known from Kü Channel, ( Sakamoto & Hayashi 1977) and Amakusa Island, Japan ( Fujino 1978); the South China Sea ( Bruce 1966, 1979, 2003; Li & Bruce 2006; Li et al. 2005) and East China Sea (Li 2006).

Reports from the Seychelle Islands, (Bruce 1976), Indonesia, Papua New Guinea and the Philippines ( Debelius 2001) are dubious. The ovigerous female Fijian specimen, from 144-150 m, referred to P. tosaensis by Li & Bruce (2006), a perfect specimen, has clearly been misidentified as the ambulatory dactyli have well developed accessory teeth. The specimen has been re-examined and is now considered to be referable to A. tenuirostris ( Bruce 1991) .

Figures

FIGURE 4. Ancylomenes tosaensis (Kubo) male, Australian Northwest Shelf, NTM Cr015053: A, anterior carapace, rostrum and bases of antennae; B, third pereiopod, propodus and dactylus; C, same, distal propodus and dactylus.