Woldstedtius bauri, Klopfstein, Seraina, 2014

Klopfstein, Seraina, 2014, Revision of the Western Palaearctic Diplazontinae (Hymenoptera, Ichneumonidae), Zootaxa 3801 (1), pp. 1-143 : 109-110

Woldstedtius bauri sp. nov.

Description. Antenna with multiporous plate sensilla present also on ventral surface of flagellomeres but often sparser than dorsally. Face very weakly elevated centrally, without vertical depressions; strongly and evenly coriaceous, at most centrally with some very indistinct punctures. Clypeus separated from face by a shallow groove, elevated basally, remainder flat or weakly concave, bilobed; coriaceous over entire surface, with some weak longitudinal striae apically. Head strongly constricted behind compound eyes. Mesoscutum strongly and evenly coriaceous and matt, with some indistinct punctures, without notauli; scutellum entirely coriaceous and matt, with lateral carinae only present basally. Mesopleuron strongly coriaceous and matt over entire surface except for a smooth area around mesopleural fovea, often more roughly coriaceous or rugose on lower than upper mesopleuron; sternaulus very weakly impressed, epicnemial carina complete also ventrally. Metapleuron coriaceous and matt. Propodeum with carinae almost completely reduced but often with front part of pleural carina and sometimes very short traces of apical part of lateral longitudinal carina present; evenly coriaceous and matt. Fore wing areolet open, vein 1cu-a usually apical of vein M; hind wing with two or three basal hamuli, vein CU+cu-a broken slightly below middle. Hind coxa coriaceous and matt. Metasoma dorsoventrally depressed; hind margins of all tergites straight or convex; tergites entirely coriaceous and matt, tergite 1 with weak longitudinal carinae on about basal half, with some longitudinal striae parallel to them; tergite 2 finely striate basally. Second and third tergites with spiracles dorsal, above lateral folds. Ovipositor sheaths straight, stout, with tip transversely truncate apically so that ovipositor tip is often protruding from it; coriaceous and with dense but rather short setae over entire surface but very concentrated around tip.

Colouration of females. Antenna black. Head and mesosoma black, face usually with a small yellow face patch which is smaller than central elevated area, clypeus black, yellow on mouthparts, hind corner of pronotum, tegula, usually subtegular ridge, and upper mesepimeron; without yellow shoulder marks, scutellum at most with a small yellow apical spot. Legs orange, fore coxa mainly yellow with black base, mid and hind coxae mainly black with a yellow apex, often with traces of reddish colour; femora orange; hind tibia black with a white base, hind tarsus dark. Metasoma black.

Colouration of males. As in females but with yellow ventrally on antenna, entire face, propleuron, and small yellow shoulder marks; yellow on epicnemium; mesosternum and lower mesopleuron variously marked with yellow, often mainly yellow. Coxae mainly yellow, hind coxa usually marked with orange and black. Metasoma black with yellow spots basally on tergites 3 and often 4

Similar species. W. bauri is similar to W. biguttatus , from which it can be distinguished by the more extensive yellow colouration and the dark coxae. It might also be confused with W. citropectoralis , the species with which it clusters in the molecular studies ( Fig. 6 ); W. bauri , however, has longer antennae, the inner orbits close to parallel and the coxae dark.

Type material. Holotype ♀ at NRM: Sweden, Småland, Nybro kommun , Bäckebo , Grytsjöns naturreservat , N56°55'17.96", E16°6'4.42", Malaise trap, leg. Swedish Malaise Trap Project, 2.–12.VII.2005. Labels: “SWEDEN Sm, Nybro; komun, Bäckebo,; Grytsjöns naturreservat,; N6310596 E1518027 (=Trap ID 1000)” [coordinates on label incorrect], “Old aspen forest in boulder terrain CollID; 1323. 2.VII.–12.VII.2005; Leg.Swedish MT Project”, “Extraction plate 4; 4-D7; label: SKlopfstein 2008”, “Holotype; Woldstedtius ♀; bauri n.sp.; det. S.Klopfstein 2010 ” . Paratypes: As holotype, 1♀, at NRM. Sweden, Småland, Gränna kommun, Lönnemalen, next to old cellar in Norway spruce forest with big harvested ashes, N58°02.935', E14°34.382', leg. SMTP, 01.VII.–17.VII.2005 . 1♀, at NRM. Sweden, Östergottland, Ödeshögs kommun, Omberg, Storpissan, N58°20.095’, E14°39.300, 28.V.–5.VII.2005 , 3♀, at NRM. Sweden, Småland, Nybro kommun, Alsterbro/Alsteran. Mixed forest, N56°56'11.53", E15°55'12.60", leg. SMTP, 01.VI.–10.VI.2006, 1♀, 05.VII.–10.VII.2005, 1♀, both at NRM.

Additional material. Sweden (13), Switzerland (2), United Kingdom (14).

Distribution. Western Palaearctic.

Figures. Habitus ( Fig. 36 A), male terminal sclerites ( Fig. 38 G).

Figures

FIGURE 6. Phylogeny of specimens of the genus Woldstedtius obtained from CO 1 (A) and ITS 2 sequences (B). The tree with the highest likelihood score is shown with clade supports obtained by 100 bootstrap iterations.

FIGURE 36. Habitus of Diplazontinae species. A. Woldstedtius bauri sp. nov. B. Woldstedtius biguttatus. C. Woldstedtius citropectoralis. D. Woldstedtius holarcticus. E. Woldstedtius patei. F. Xestopelta gracillima.

FIGURE 38. Ninth sternite and ninth and tenth tergites of male Diplazontinae species. A. Sussaba erigator. B. Sussaba pulchella. C. Sussaba roberti. D. Syrphophilus asperatus. E. Tymmophorus erythrozonus. F. Tymmophorus obscuripes. G. Woldstedtius bauri. H. Woldstedtius citropectoralis. I. Woldstedtius holarcticus. J. Xestopelta gracillima.