Gracilipsodes lanceolatus, Malm & Johanson, 2008

Malm, Tobias & Johanson, Kjell A., 2008, Revision of the New Caledonian endemic genus Gracilipsodes (Trichoptera: Leptoceridae: Grumichellini), Zoological Journal of the Linnean Society 153 (3), pp. 425-452 : 438-440

GRACILIPSODES LANCEOLATUS SP. NOV. ( FIGS 7 , 13 )

Diagnosis: This species is separated from the other Gracilipsodes species by having the tibial spur formula 2, 2, 1, and by having only fork 5 present in the hindwings. The genitalia are unique in having a complete tergum X lacking pre-apical lateral processes.

Description, male: Body medium brown. Wings greyish brown with blackish hairs. Tibial spur formula 2, 2, 1. Hind tibiae each with 20–26 small blackish spines (N = 10). Forewing ( Fig. 7A ): length 5.7–6.1 mm (N = 10); narrow, apically rounded; forks 1 and 5 present; crossvein m–cu nearly as long as crossvein r–m, forms a wide angle with M 3+4; crossvein r–m at sharp angle to M. Hindwing ( Fig. 7B ): length 3.9– 4.2 mm (N = 10); slender, apex nearly acute; fork 5 present; crossvein r–m straight; costa with 22 hamuli.

Genitalia: Segment IX annular, laterally setose, widest at mid height; lateral part of tergite IX as wide as sternite IX ( Fig. 7C ); posterior margins of sternum IX and tergum IX produced posteriorly ( Fig. 7C, D ), lateral margins of sternum IX produced anteriad ( Fig. 7E ). Superior appendages setose, flattened dorsoventrally, slightly longer than tergum X ( Fig. 7C ), apices rounded in dorsal view ( Fig. 7D ). Tergum X entire, membranous, tubular; basally and apically wide, apex rounded, dorsal margin straight in lateral view ( Fig. 7C ), narrowing apically, apex with notch in dorsal and ventral views ( Fig. 7D, E ); pre-apical lateral processes absent. Inferior appendages each bipartite, setose; basal part slender at base in lateral view ( Fig. 7C ), mesally concave, produced posteromesally into slender, tapering, and apically rounded lip, bearing small dorsal spines ( Fig. 7C, E ); apical part of each inferior appendage digitate, slender, curving mesally; in lateral view, curving slightly dorsally, widening slowly towards rounded apex, with 25 mesal spines ( Fig. 7C ); mesal process with four spines ( Fig. 7D ). Harpago reduced. Phallic apparatus simple ( Fig. 7F, G ); phallicata basally tubular, fused with phallobase, apex truncate; curving slightly ventrally along its length; apex with pair of pre-apical, sclerotized, ventrally produced broad processes, being directed posterolaterally, and with rounded apices ( Fig. 7F ); phallotremal sclerite U-shaped ( Fig. 7G ).

Holotype (male): New Caledonia, Province Sud, western slope of Mt Ningua, Kwé Néco Stream, at Camp Jacob , 3.7 km west-northwest of the summit of Mt Ningua , on Bouloparis-Thio Road , about 50 m upstream of road, 21°43.613′S, 166°06.567′E, 150 m a.s.l., 29 November–12 December 2003, Malaise trap, loc#054 (Johanson).

Paratypes: Four males, same data as holotype ; one male, ditto, except 12 December 2003 – 5 January 2004 ; 30 males, New Caledonia, Province Sud, western part of Mt Ningua, Kwé Néco Stream, 3.9 km west of summit of Mt Ningua , on Bouloparis-Thio Road , about 50 m upstream of road, 21°44.359′S, 166°06.009′E, 117 m a.s.l., 20 November 2003 – 12 December 2003, Malaise trap, loc#035 (Johanson) ; one male, New Caledonia, Province Sud, western slope of Mt Ningua, Kwé Néco Stream, at Camp Jacob , 3.9 km west of summit of Mt Ningua , on Bouloparis-Thio Road , about 50 m upstream of road, 21°44.083′S, 166°06.298′E, 117 m a.s.l., 29 November 2003 – 12 December 2003, Malaise trap, loc#053 (Johanson) .

Distribution: Province Sud.

Etymology: lanceolatus , from Latin, lancea, spearlike; named after the slender and apically acute hindwings.

Figures

Figure 7. Gracilipsodes lanceolatus sp. nov. A, forewing. B, hindwing. C, genitalia, lateral. D, genitalia, dorsal. E, genitalia, ventral. F, phallus, lateral. G, phallus, ventral.

Figure 13. Maps of New Caledonian Grande Terre with plotted distributions of the different species. A, Gracilipsodes aoupiniensis sp. nov. (triangles), Gracilipsodes aureus sp. nov. (white circles), Gracilipsodes grandis sp. nov. (squares), Gracilipsodes koghiensis sp. nov. (diamond), Gracilipsodes lanceolatus sp. nov. (black circle). B, Gracilipsodes aurorus sp. nov. (triangle), Gracilipsodes robustus sp. nov. (black diamond), Gracilipsodes psocopterus Sykora, 1967 (circles), Gracilipsodes similis Ward (2001) (squares).