Eupines (Byraxis) lewisi Broun, 1910

Shen, Jia-Wei & Leschen, Richard A. B., 2020, Revision of Eupines King of New Zealand (Coleoptera: Staphylinidae: Pselaphinae: Goniaceritae), Zootaxa 4777 (1), pp. 1-84 : 46-47

Eupines (Byraxis) lewisi Broun, 1910

Fig. 29 ; map Fig. 52

Eupines lewisi Broun, 1910: 24 . Raffray 1911: 83, 1924: 157; Hudson 1923: 364 and 367, 1934: 183 and 185.

Eupines (Byraxis) lewisi : subgeneric assignment by Nomura and Leschen 2006: 255. Maddison 2010: 417; Shen and Leschen 2019: 268 .

Type locality: [ MC] Broken River , Canterbury .

Type depository: BMNH HT ♂

Diagnosis: Head with vertexal foveae punctiform; antennae 10-segmented with 2-segmented antennal club; A4, A6, A7, A8 and A10 unmodified, A5 large and asymmetrical, bean-shaped, lateral margin laminate and curved, forming bowl-shaped excavation, A7 asymmetrical, A8 disc-shaped, A9 large and asymmetrical, slightly shorter than A10, inner side strongly excavated in dorsal view, with upper margin laminate and extended, forming gulfshaped structure, A10 largest and conical, about 1.2 × longer than A9. Abdominal ventrite 2 with pair of large discal protuberances slightly curving towards each other, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Protrochanters with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified.

Redescription: Body length 1.58–1.60 mm; body colour dark brown with lighter coloured legs. Head about as long as wide, frons slightly narrowed at end, about half width of head at eye level; eyes moderately developed, with about 29 facets; vertexal foveae punctiform; antennal tubercles weak; lacking postantennal notches; postantennal cavities medium-sized, about half length of antennal tubercles in dorsal view; antennae 10-segmented with 2-segmented antennal club; A4, A6, A7, A8 and A10 unmodified, A5 large and asymmetrical, bean-shaped, lateral margin laminate and curved, forming bowl-shaped excavation, A7 asymmetrical, A8 disc-shaped, A9 large and asymmetrical, slightly shorter than A10, inner side strongly excavated in dorsal view, with upper margin laminate and extended, forming gulf-shaped structure, A10 largest and conical, about 1.2 × longer than A9. Pronotum about as long as wide, wider than head at eye level; lacking lateral antebasal foveae. Elytra as long as wide, widest at apical two-thirds; humeral calli distinctively prominent; sutural stria distinctively extending from base to apex. Abdomen with tergite 1 about 1.8 × longer than tergite 2; ventrite 2 with pair of large discal protuberances slightly curving towards each other, inserted with strongly curved small apical setae; ventrite 6 with pair of short and acute median struts, about one-fourth of ventral length. Legs with protrochanters modified with small ventral spine, about one-third of protrochanteral width; protibiae and mesotibiae unmodified. Aedeagus about 0.32 mm long, with sinuate parameres long and stout, widest near basal one-fourth, apex strongly bent downwards and twisted in lateral view, with three short apical setae; median lobe with two short sclerites, much shorter and narrower than parameres in dorsal view, left one distinctively curved outwards, right one point inwards.

Comments: The special antennal modifications ( Figs. 29 B–D) can easily separate this species from all congeners.

Distribution: North Island. BP. South Island. NN, MC.

Additional material examined: 5 ♂♂, BP : 2, BMNH, 3052. ♂, Rotoiti , 10/3/16, New Zealand, Broun Col- lection. Brit. Mus., 1922–482; NN: 2, FMNH , Nelson Lakes N.P., N. slope Mt. Robert, Pinchgut Track , 950 m, 14/ XII/84–6/I/85, Nothofagus spp. forest, A. Newton /M. Thayer, 707, flight intercept (window) trap; MC: 1, BMNH , 3052, ♂, Rakaia , 9/8/1912, New Zealand, Broun Collection. Brit. Mus., 1922–482 .

Figures

FIGURE 29. Diagnostic characters of E. (B.) lewisi Broun, 1910. A) Habitus. B) Antenna, in dorsal view. C) Same, in lateral view. D) Same, in ventral view. E) Protrochanter. F) Ventrite 2. G) Ventrite 6. H) Aedeagus, in dorsal view. I) Same, in lateral view. J) Same, in ventral view. Scale bars: A = 1 mm, B–F = 0.2 mm, G–J = 0.1 mm.

FIGURE 52. Geographic distributions of E. (B.) impar (Sharp, 1874), E. (B.) impressifrons (Broun, 1880), E. (B.) insolita sp. n., E. (B.) lewisi Broun, 1910, E. (B.) longiceps Raffray, 1904, E. (B.) mayae sp. n., and E. (B.) micans (Sharp, 1874) in New Zealand.