Kirbyana Distant, 1906

Zhi, Yan, Yang, Lin & Chen, Xiang-Sheng, 2021, Two new bamboo-feeding species of the genus Kirbyana Distant, 1906 from China (Hemiptera, Fulgoromorpha, Cixiidae), ZooKeys 1037, pp. 1-14 : 1

Genus Kirbyana Distant, 1906

Kirbya Melichar, 1903: 37, preoccupied by Kirbya Robineau-Desvoidy, 1830 ( Diptera ).

Kirbyana Distant, 1906a: 262, nom. nov. for Kirbya Melichar, 1903.

Kirbyella Kirkaldy, 1906: 248, synonymised by Distant 1906b: 274.

Saccharias Kirkaldy, 1907: 125, synonymised by Fennah 1980: 239.

Commolenda Distant, 1911: 741, synonymised by Fennah 1978: 211.

Type species.

Kirbya pagana Melichar, 1903 (original designation by Distant).

Diagnosis.

Head. Head including eyes slightly narrower than pronotum. Vertex in profile horizontal, in same line as thorax, meeting frons abruptly nearly at 90°; subapical carina absent. Frons somewhat longer than broad or as long as wide; median carina present; anterior margin angularly emarginate or transverse; position of maximum width of frons more or less at the level of antennae. Median ocellus absent. Subapical segment of rostrum 2.5 times longer than apical segment.

Thorax. Pronotum very narrow, scarcely half as long as vertex in midline. Mesonotum nearly twice as long as pronotum and vertex together. Forewing with the eucarpian basal concavity of the anterior margin of the forewings, also slightly concave at node level. Hindwing with simple radius (R). Hind tibia lacking lateral spines. Metatibiotarsal formula: 6/8-11/9-11.

Male genitalia. Pygofer symmetrical and prolonged with symmetrical lateral lobes in lateral view. Medioventral process thumb-like in lateral view. Anal segment short and stout. Gonostyli relatively small and symmetrical. Aedeagus slender and endosoma of aedeagus with spinose processes.

Female genitalia. Ovipositor elongate, orthopteroid and slightly curved upwards; anal segment square or rectangular in dorsal view; 9th tergite without wax plate. Posterior vagina with sclerites.

Distribution.

Australian and Oriental regions.

Checklist and distributions of species of Kirbyana Distant

K. aspina Zhi & Chen, sp. nov.; China (Hunan).

K. australis (Muir, 1913); Australia (Northern Territory; Queensland).

K deusta (Distant, 1911); Central India.

K deventeri (Kirkaldy, 1907); Indonesia (Java).

K. furcata Zhi & Chen, sp. nov.; China (Guangxi, Yunnan).

K javana Muir, 1913; Indonesia (Java).

K lini Tsaur & Hsu, 2003; China (Taiwan).

K pacifica Emeljanov & Hayashi, 2007; China (Taiwan), Japan (Ryukyu Islands).

K. pagana (Melichar, 1903); Sri Lanka (Peradeniya).

K pratti Muir, 1913; Malaysia (Parit Buntar).

K pratti thyas Fennah, 1978; Vietnam (Cuc-phuong Province, Ninh Binh).

Key to the species (males) of Kirbyana Distant

1	Anterior margin of vertex transverse	2
-	Anterior margin of vertex angularly excavated or incised at middle	7
2	Forewing with numerous small fuscous spots on basal half and several waved fuscous lines on apical half	K deusta (Distant, 1911)
-	Forewing not so marked	3
3	Forewing with a transverse veinlet from M to CuA near Cu fork	4
-	Forewing without transverse veinlet from M to CuA near Cu fork	5
4	Forewing without transverse veinlet from MP3+4 to CuA1 ( Löcker et al. 2010: fig. 6C)	K. australis (Muir, 1913)
-	Forewing with two transverse veinlets from MP3+4 to CuA1 ( Kirkaldy 1907: fig. 1)	K deventeri (Kirkaldy, 1907)
5	Ventral margin of periandrium without spinous processes near base ( Tsaur and Hsu 2003: fig. 2D)	K lini Tsaur & Hsu, 2003
-	Ventral margin of periandrium with two spinous processes near base	6
6	One of the two basal processes of ventral margin of periandrium bifurcate (Fig. 5I, J, L )	K. furcata sp. nov.
-	Both basal processes of ventral margin of periandrium unbifurcated (Fig. 2I, J, L )	K. aspina sp. nov.
7	All tubercles of longitudinal veins in forewings colourless	K javana Muir, 1913
-	Some tubercles of longitudinal veins in forewings dark	8
8	Forewing with MP joining CuA with crossvein M3+4-CuA ( Emeljanov and Hayashi 2007: fig. 25)	K pacifica Emeljanov & Hayashi, 2007
-	Forewing with MP joining CuA directly without crossvein M3+4-CuA	9
9	Forewing with a series of black spots on the claval vein and the inner bifurcating veins; mesonotum with a small dark spot near base of each lateral carina	K. pagana (Melichar, 1903)
-	Forewing not so marked; mesonotum without a spot on each side near base of lateral carina	K pratti Muir, 1913

Figures

Figure 2. Kirbyana aspina sp. nov., male A head and thorax, dorsal view B face, ventral view C forewing D apex of left hind leg, ventral view E genitalia, lateral view F pygofer and gonostyli, ventral view G anal segment, dorsal view H gonostyli, inner lateral view I aedeagus, right side J aedeagus, left side K aedeagus, dorsal view L aedeagus, ventral view. Scale bars: 0.5 mm (A, B, E-L); 1.0 mm (C).

Figure 5. Kirbyana furcata sp. nov., male A head and thorax, dorsal view B face, ventral view C forewing D apex of left hind leg, ventral view E genitalia, lateral view F pygofer and gonostyli, ventral view G anal segment, dorsal view H gonostyli, inner lateral view I aedeagus, right side J aedeagus, left side K aedeagus, dorsal view L aedeagus, ventral view. Scale bars: 0.5 mm (A, B, E-L); 1.0 mm (C).