Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Genomics-based taxonomic rearrangement of Achlyodini and Carcharodini (Lepidoptera: Hesperiidae: Pyrginae), Insecta Mundi 2023 (16), pp. 1-33 : 15-16

Thilla Grishin , new subgenus

http://zoobank.org/ ADF9A522-9D1D-466B-8A62-65A40F9E70F2

Type species. Eurypterus later Mabille, 1891 .

Definition. The genomic tree shows that the genus Eantis Boisduval, 1836 (type species Urbanus thraso Hübner, 1807 ) splits into five lineages at the subgenus level, and we regard these lineages as representing five subgenera ( Fig. 3 ). Out of these subgenera, in addition to the nominotypical, only one has a name: Tosta Evans, 1953 (type species Tosta tosta Evans, 1953 ), while three others are new. One of these new subgenera consists of species previously placed in Aethilla Hewitson, 1868 (type species Aethilla eleusinia Hewitson, 1868 ) and keys to F.1.8 or F.1.4a (in part, as incorrect synonymy) in Evans (1953) and is distinguished from its relatives by the following characters: forewing more pointed or even slightly falcate at the apex, hindwing less produced or more rounded at the tornus; ampulla with a long terminally serrated process directed dorsoposterad and extending over harpe, harpe with process-like expansion at its base by ampulla. In DNA, a combination of the following nuclear genomic base pairs is diagnostic: aly577.14.5:A210G, aly728.5.4:G78A, aly461.13.3:T129C, aly2012.59.3:A90G, aly2012.59.3:T100C.

Etymology. The name is a feminine noun in the nominative singular, formed from the previous genus name for these species: [Ae] Thilla.

Species included. Eurypterus later Mabille, 1891 and Eurypterus haber Mabille, 1891 .

Parent taxon. Genus Eantis Boisduval, 1836 .

Figures

Figure 3. The nuclear genome-based phylogeny of Achlyodini (continued from Fig. 1). A segment corresponding to the subtribe Achlyodinia is shown. The top segment of the tree (subtribe Pythonidina) is replaced with four dots and is shown in Fig. 1. See Fig. 1 for notations. The names of taxa that before this work were placed in the genus Aethilla are shown in orange to illustrate the lack of monophyly.